Open Journal of Archaeometry 2014; volume 2:5306
[Open Journal of Archaeometry 2014; volume 2:5306] [page 81]
Exploring the use of stable
carbon isotope ratios in
short-lived leporids for local
paleoecological reconstruction
Stephen Smith,
1
Raymond Mauldin,
2
Cynthia M. Munoz,
2
Robert Hard,
1
Debajyoti Paul,
3
Grzegorz Skrzypek,
4
Patricio Villanueva,
5
Leonard Kemp
6
1
Department of Anthropology, University
of Texas at San Antonio;
2
Center for
Archaeological Research, University of
Texas at San Antonio, TX, USA;
3
Department of Civil Engineering
(Geosciences), Indian Institute of
Technology, Kanpur, India;
4
West Australian Biogeochemistry
Centre, School of Plant Biology, The
University of Western Australia, Crawley,
Australia;
5
Department of Geology,
University of Texas at San Antonio, TX;
6
Geo-Marine, El Paso, TX, USA
Abstract
Most ecological proxies used in archaeolog-
ical research operate at scales that are too
coarse-grained for consideration of hunter-
gatherer adaptive decisions. Hunter-gatherers
adapt to local ecological conditions and short
(e.g. seasonal, yearly) time frames. Our goal is
to develop proxies to identify ecological shifts
at fine-grained temporal and spatial scales for
archaeological research. We use stable carbon
isotope ratios (δ
13
C) in bone collagen from 58
modern leporids from two distinct ecological
areas in the American Southwest as a proxy to
reconstruct vegetation and climate patterns at
fine-grained scales. Higher δ
13
C values in col-
lagen of cottontail (Sylvilagus sp.) and jackrab-
bit (Lepus sp.) collected in the northern
Chihuahuan Desert of New Mexico and West
Texas suggest a more C
4
/crassulacean acid
metabolism (CAM) dominated local vegetation
when contrasted to samples from Kerr County
in Central Texas. Comparisons of temperature
and precipitation patterns between the two
areas, along with vegetation data, suggest that
key ecological differences, reflected in the col-
lagen isotopic compositions, are likely related
to rainfall amounts and the type of green for-
age available to leporids, especially during
winter months. Leporids in dry areas may be
relying on CAM plants, including prickly pear,
which has a C
4
isotopic signature. Alternative
resources are likely to be available in wetter
areas such as Central Texas.
Introduction
Reconstructions of past ecological condi-
tions in archaeological research often rely on
proxy data, such as shifts in pollen frequen-
cies, which operate at variable spatial and tem-
poral scales. While appropriate for many con-
cerns, most ecological proxies operate at
scales that are too coarse-grained for consider-
ation of human adaptive decisions. Hunter-
gatherers and agriculturalists adapt to local
rather than regional ecological conditions, and
to changes that occur on a seasonal or yearly
time frame, not shifts that happen over cen-
turies (Winterhalder and Leslie, 2002). Arch-
aeologists need to develop high-resolution eco-
logical proxies that operate at scales that are
coterminous with those used by humans in
making adaptive decisions.
In this paper, we explore the use of stable
carbon isotope ratios (δ
13
C) of bone collagen
in cottontails (Sylvilagus sp.) and jackrabbits
(Lepus sp.) as potential high-resolution prox-
ies to reconstruct vegetation and climate pat-
terns at short-term temporal and spatial scales.
We have used samples from an archaeological
assemblage from Texas, USA to monitor eco-
logical shifts (Munoz et al., 2011; Kemp, 2008)
and are currently investigating other archaeo-
logical assemblages in the American
Southwest. We focus on refining the applica-
tion through a comparison of δ
13
C values in
bone collagen (δ
13
C
collagen
) of leporids from two
distinct ecological settings. δ
13
C values of mod-
ern leporids collected in southern New Mexico
and West Texas (n=26) suggest that these
areas have a stronger C
4
/CAM signature rela-
tive to those collected in Kerr County in
Central Texas (n=32). Comparisons of temper-
ature and precipitation patterns between the
two areas, along with vegetation transects
data, suggest that key ecological differences
reflected in the collagen are likely related to
rainfall amounts and the type of green forage
available to leporids, especially during the win-
ter months.
As a paleoecological proxy, leoprids have
several advantages. Jackrabbit and cottontail
diets incorporate a variety of plants, with
grasses, forbs, shrubs, and succulents con-
sumed in response to availability, growing con-
ditions (i.e. temperature, rainfall, and plant
dormancy), and moisture content. Analyses of
the stomach contents of Lepus collected in
southern Arizona (Vorhies and Taylor, 1933)
suggest a seasonal preference for grasses in
the spring and midsummer, following rains. As
grass production declines in early fall and win-
ter, rabbits concentrate on browse consisting
primarily of mesquite (Prosopis glandulosa).
Cactus, such as prickly pear (Opuntia sp.) and
cholla (Echinocereus fendleir), are more heav-
ily consumed during drier months. Similar
seasonal trends, with grasses preferred in the
late spring, summer, and early fall, forbs and
shrubs in the fall and winter, and shrubs and
succulents in winter and early spring are
reported for jackrabbits in a variety of regions
(Best, 1996; Currie and Goodwin, 1966; Fatehi
et al., 1988; Hayden, 1966; Hoffman et al., 1993;
Correspondence: Cynthia M. Munoz, Center for
Archaeological Research, 1 UTSA Cir, University
of Texas at San Antonio, San Antonio, 78249 TX,
USA.
Tel./Fax: +1.210.458.4397.
E-mail: cindy.munoz@utsa.edu
Key words: carbon isotopes, paleoecological
reconstruction, leporids.
Acknowledgements: thanks to Richard Doucett
and Melanie Caron [Colorado Plateau Analytical
Laboratory, Nothern Arizona University (NAU),
Flagstaff, AZ], to Bruce Moses who drafted the
figures for this manuscript, and to the Center for
Archaeological Research - The University of
Texas at Sant Antonio (CAR-UTSA) staff. Kevin
Daiber, Kelly Denham, and Lynn Wack of CAR
helped with sample preparation. Thanks to Heath
Garner of the Texas Tech University Museum
(Lubbock) and Dr. Robert Baker, National
Science Research Laboratory, at Texas Tech for
graciously allowing us to sample their modern
faunal collections. Bryant Saner provided addi-
tional modern specimens. Thanks to Dr. Mark
Pollard (Archaeological Science Research Lab for
Archaeology, University of Oxford) for reviewing
this manuscript and to Robert Tykot [Department
of Anthropology, University of South Florida
(USF), Tampa, FL] for compiling and editing the
Proceeding of the 2010 ISA. This research is sup-
ported by NSF grant #0804169 awarded to
Raymond Mauldin and Robert Hard. All data sets
used here, as well as the samples themselves in
many cases, are available through
Raymond.mauldin@utsa.edu. Details on the
research can be found at http://car.utsa.edu
Citation: Smith S, Mauldin R, Munoz CM, Hard R,
Paul D, Skrzypek G, Villanueva P, Kemp L, 2014.
Exploring the use of stable carbon isotope ratios
in short-lived leporids for local paleoecological
reconstruction. In: RH Tykot (ed.), Proceedings of
the 38
th
International Symposium on
Archaeometry – May 10
th
-14
th
2010, Tampa,
Florida. Open Journal of Archaeometry 2:5306.
Presented at the 38
th
International Symposium on
Archaeometry – May 10
th
-14
th
2010, Tampa,
Florida.
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Licensee PAGEPress, Italy
Open Journal of Archaeometry 2014; 2:5306
doi:10.4081/arc.2014.5306
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