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J Comp Physiol B
DOI 10.1007/s00360-014-0835-y
ORIGINAL PAPER
Thermoregulatory plasticity in free-ranging vervet monkeys,
Chlorocebus pygerythrus
Alwyn Lubbe · Robyn S. Hetem · Richard McFarland ·
Louise Barrett · Peter S. Henzi · Duncan Mitchell ·
Leith C. R. Meyer · Shane K. Maloney · Andrea Fuller
Received: 26 February 2014 / Revised: 12 May 2014 / Accepted: 28 May 2014
© Springer-Verlag Berlin Heidelberg 2014
inactive. These low minimum body temperatures were
associated with low black globe temperature (GLMM,
β = 0.046, P < 0.001), short photoperiod (β = 0.010,
P < 0.001) and low rainfall over the previous 2 months,
which we used as a proxy for food availability (β = 0.001,
P < 0.001). Despite the lower average winter minimum
body temperatures, there was no change in the lower modal
body temperature between winter and summer. Therefore,
unlike the regulated physiological adjustments proposed
for torpor or hibernation, these minimum winter body tem-
peratures did not appear to reflect a regulated reduction in
body temperature. The thermoregulatory plasticity never-
theless may have fitness benefits for vervet monkeys.
Keywords Bio-logging · Climate change ·
Heterothermy · Primate · Body temperature
Introduction
In seasonally-variable habitats, mammals are faced with
the challenge of maintaining body temperature while being
exposed to dynamic thermal environments and contend-
ing with fluctuations in food and water availability. At low
environmental temperatures, typically in conjunction with
food scarcity, some mammals—termed heterotherms—
employ physiological mechanisms such as torpor or hiber-
nation (i.e., prolonged torpor) that result in a decrease in
energy expenditure (Geiser 2004). Torpor is defined as
a state of inactivity and reduced responsiveness (IUPS
Thermal Commission 2001), typically associated with a
temporary, but substantial, reduction in body temperature
below body temperature thresholds of about 35 °C (Willis
2007) to 30 °C (Barclay et al. 2001). However, recent stud-
ies have revealed significant seasonal and daily variation
Abstract We used implanted miniature data loggers to
obtain the first measurements of body temperature from a
free-ranging anthropoid primate. Vervet monkeys (Chlo-
rocebus pygerythrus) living in a highly seasonal, semi-
arid environment maintained a lower mean 24-h body
temperature in winter (34.6 ± 0.5 °C) than in summer
(36.2 ± 0.1 °C), and demonstrated increased heterothermy
(as indexed by the 24-h amplitude of their body tempera-
ture rhythm) in response to proximal environmental stress-
ors. The mean 24-h amplitude of the body temperature
rhythm in summer (2.5 ± 0.1 °C) was lower than that in
winter (3.2 ± 0.4 °C), with the highest amplitude for an
individual monkey (5.6 °C) recorded in winter. The higher
amplitude of the body temperature rhythm in winter was a
consequence primarily of lower 24-h minimum body tem-
peratures during the nocturnal phase, when monkeys were
Communicated by I. D. Hume.
A. Lubbe (*) · R. S. Hetem · R. McFarland · L. Barrett ·
D. Mitchell · L. C. R. Meyer · S. K. Maloney · A. Fuller
Brain Function Research Group, Faculty of Health Sciences,
School of Physiology, University of the Witwatersrand,
7 York Road, Medical School, Parktown, Johannesburg 2193,
South Africa
e-mail: alwynlubbe@gmail.com
L. Barrett · P. S. Henzi
Department of Psychology, University of Lethbridge,
Lethbridge, Canada
P. S. Henzi
Applied Behavioural Ecology and Ecosystem Research Unit,
University of South Africa, Johannesburg, South Africa
S. K. Maloney
School of Anatomy, Physiology and Human Biology,
University of Western Australia, Crawley, Australia