The wild goats Capra aegagrus Erxleben, 1777 of the
Mediterranean Sea and the Eastern Atlantic Ocean islands
MARCO MASSETI
Laboratories of Anthropology and Ethnology, Department of Evolutionistic Biology,
University of Florence, Via del Proconsolo, 12 – 50122 Florence, Italy
ABSTRACT
1. Populations of wild goats that can be referred to as phenotypes of the pasang, or Bezoar
goat, or wild goat Capra aegagrus Erxleben, 1777, still occur on several of the Mediterranean
and Eastern Atlantic islands. Other populations became extinct not many decades ago.
2. Fossil evidence for the natural spread of the wild goat to any of these islands has not been
found. Originating in the Near East, the region of its natural range and its earliest domesti-
cation, the species was introduced by humans onto the Mediterranean islands starting as
early as the Pre-Pottery Neolithic.
3. The islands of the Eastern Atlantic were colonized artificially by animals of ‘pre-Hispanic’
origin. The principal phenotypes expressed by all these populations of goats can be matched
with the characteristics of both C. a. aegagrus Erxleben, 1777 (aegagrus phenotype) and
C. a. dorcas Reichenow, 1888 (dorcas phenotype).
4. Most of these animals currently survive on the islands without being fed by humans, and
should be protected.
Keywords: Capra aegagrus, domestic animals, Eastern Atlantic islands, feral animals,
Mediterranean islands, wild goat
Mammal Review (2009), 39, 141–157
doi: 10.1111/j.1365-2907.2009.00141.x
INTRODUCTION
Populations of wild goats Capra aegagrus Erxleben, 1777 have been present on several of the
Mediterranean and Eastern Atlantic islands from time immemorial. It used to be thought
that they were autochthonous forms. However, in view of the lack of fossil evidence for these
ungulates in mainland Europe or on the islands, they are now considered ancient feral breeds
deriving from human importations in ancient or more recent times (Payne, 1968; Clutton-
Brock, 1981; Masseti, 1981; Davis, 1984, 1987; Helmer, 1989; Ciani & Masseti, 1991). The
wild goat is in fact completely absent from the original late Pleistocene fauna of southern
Europe (Schultze-Westrum, 1963; Azzaroli, 1983), its original range being documented only
from the north-eastern Mediterranean mainland, where wild goat populations still survive in
the south-western and eastern Anatolian mountains. They are found in inner south-western
Asia, from south-western Anatolia to the western Indian subcontinent, throughout the
eastern Caucasus, southern Turkmenistan, Afghanistan and south-western Pakistan (Harri-
son, 1968; Harrison & Bates, 1991; Masseti, 1997a; Pedrotti & Lovari, 1999; Grubb, 2005). In
the Near East, the altitudinal range of the species extends from sea level to the alpine zone.
The goats are capable of surviving in very arid regions where the vegetation is sparse; their
Correspondence: M. Masseti. E-mail: marco.masseti@unifi.it
Mammal Rev. 2009, Volume 39, No. 2, 141–157. Printed in Singapore.
© 2009 The Author. Journal compilation © 2009 Mammal Society, Mammal Review, 39, 141–157