Reference : Biol. Bull., 149 : 287—315. (October, 1975)
BIOTIC CENSUS OF CAPE COD BAY :
DALE R CALDER
Marine Resources Research Institute, P. 0. Box 12559, Charleston, South Carolina 29412
The Biotic Census of Cape Cod Bay was initiated in 1965 by the Systematics
Ecology Program at the Marine Biological Laboratory, Woods Hole, Massachusetts.
Completed in 1969, the census involved qualitative and quantitative sampling of
the benthos to determine the species occurring in the bay, and to provide informa
tion about their ecology. This report is an account of the hydroids collected during
the project.
The hydroids of the northeast have received greater attention than those from
any other geographic area of North America. Over 75 papers bearing on the
taxonomy and distribution of hydroids from New England and Atlantic Canada
have been published since Stimpson's (1854) synopsis of the marine invertebrates
of Grand Manan Island. Among those not already cited in Fraser's (1944)
monograph are papers by Ruebush (1939), Crowell (1945, 1947), Fraser (1945),
Berrill (1948), Merrill (1967), Bush and Zinn (1970), Templeman (1973), and
Calder (1974), in which one or two species have been discussed. Others, including
Préfontaine (1932), Prat (1933), Procter (1933), Dexter (1944, 1947), Fuller
(1946), Woods Hole Oceanographic Institution (1952), Bousfield and Leim
(1960), Brunel (1962, 1963, 1970), Pr@fontaine and Brunel (1962), Caddy
(1970), and Bourget and Lacroix (1973), have included hydroids in a general
biotic survey or in a study on fouling organisms. Vervoort (1972) included
several species of hydroids from the northeast collected during cruises from the
Lamont-Doherty Geological Observatory. A number of hydroid species from
Newfoundland and Anticosti Island have been described by Leloup (1939, 1960),
and papers on the hydroids of northern Canada by Calder (1970, 1972) included
a number of species from the Strait of Belle Isle. Further sources of information
on hydroids of the northeast include Fraser's (1946) book on distribution and rela
tionship in American hydroids, and the identification manuals of Miner (1950),
Smith (1964), and Gosner (1971). Despite this volume of work, little specific
information is available concerning the hydroids of Cape Cod Bay.
The collection of hydroids from the biotic census was relatively small; Cape
Cod Bay apparently does not have an especially rich hydroid fauna. Nevertheless,
a number of interesting hydroids were present in the collection, particularly those
species adapted to substrates of mud and sand.
The systematic arrangement used in this report differs significantly from that
employed by Fraser (1944). While Fraser's contributions to North American
hydroid taxonomy and zoogeography remain of monumental value, the system of
classification in his 1944 monograph is little advanced over that used 75 years
earlier. Unfortunately, no comprehensive treatise on the hydroids of North
1 Contribution No. 44 from tbe South Carolina Marine Resources Center, Charleston,
South Carolina 29412.
287