Revision of the rudist genus Orestella Lupu, 1982 (Bivalvia, Order
Hippuritida) from the Upper Cretaceous of Romania
Sacit
€
Ozer
a, *
, Liana S
as
aran
b
a
Dokuz Eylül University, Engineering Faculty, Department of Geological Engineering, Tınaztepe Campus, Buca, TR-35160
_
Izmir, Turkey
b
Babes ¸-Bolyai University, Department of Geology, 1 Mihail Kog alniceanu Street, 400084 Cluj-Napoca, Romania
article info
Article history:
Received 22 June 2013
Accepted in revised form 3 September 2014
Available online
Keywords:
Rudist
Bivalvia
Orestella
Revision
Upper Cretaceous
Romania
abstract
Re-examination of the Orestella Lupu, 1982 (formerly Orestia Lupu, 1972) type material in the Geological
Institute, Bucharest, Romania, revealed the need for revision of its taxonomic status. The holotype and
paratypes of this taxon show the diagnostic characteristics of the Family Hippuritidae, not the Radio-
litidae as previously indicated. This genus should therefore be transferred to the Hippuritidae. The
structure of the pillars, the ligamental ridge and the outer shell layer of the right valve as illustrated by
the type material point to an affiliation either with Hippurites or with Hippuritella. We discuss this
assignment by taking into account the similarities with Hippurites organisans (de Montfort), Hippuritella
lapeirousei Goldfuss and Hippuritella variabilis (Munier-Chalmas). The stratigraphic framework of the
study material is also discussed with respect to the Upper Cretaceous successions in the Central-Eastern
Carpathians of Romania.
© 2014 Elsevier Ltd. All rights reserved.
1. Introduction
The new rudist genus Orestia was described and assigned to the
Radiolitidae by Lupu (1972) based on her interpretation of the outer
shell layer of the right valve in five specimens recorded by Mir aut ¸ a
and Mir aut ¸ a (1964) from the bioclastic breccia of Campa-
nianeMaastrichtian age (Lower Hangu Beds) from the Cuejdiu
Valley, north of the town of Piatra Neamt in the central area of the
East Carpathians (Fig. 1). Later, Lupu (1982) replaced the genus
name by Orestella due to preoccupation of the former name by
Orestia Chevrolat in Dejean, 1836, p. 440 (Insecta, Order Coleop-
tera). Subsequently, no additional specimens of this taxon were
recorded either in the Romanian Upper Cretaceous rudist-bearing
formations or in the Mediterranean Tethys area in general (cf.
Steuber, 2002). Nevertheless, the assignment of Orestella to the
Family Radiolitidae was carried over in the classification of rudists
proposed for the revised Bivalvia volumes of the ‘Treatise on
Invertebrate Paleontology’ (Carter et al., 2011), and also more
recently by Skelton (2013).
The main aim of our study is the taxonomic revision of Orestella
Lupu, 1982 based on re-investigation of the holotype and paratypes
housed in the Geological Institute of Romania collection in
Bucharest. The age and stratigraphic assignment of the study ma-
terial are also discussed. A new taxonomic status of Orestella is
proposed for inclusion in the revision of the Bivalvia volumes of the
‘Treatise on Invertebrate Paleontology’.
2. Geological setting
The specimens were collected by Mir aut ¸ a and Mir aut ¸ a (1964)
from the Cuejdiu Valley, north of Piatra-Neamt ¸ . Stratigraphically,
the deposits belong to the Leps ¸ a Formation from the Vrancea
Nappe (Marginal Folds Nappe, sensu S andulescu, 1984) represent-
ing the external tectonic units of the Outer Moldavidian domain of
the Romanian Carpathians (Fig. 1). The sedimentary succession of
the Vrancea Nappe (Marginal Folds) range in age from the Early
Cretaceous to the Early Miocene (Dumitrescu, 1952; B ancil a, 1958;
Grasu et al., 1988; Guerrera et al., 2012) and the following units crop
out in ascending stratigraphic order: the S arata, Leps ¸ a, Piatra
Uscat a, Jgheabu Mare, Doamna Limestone, Bisericani, Globigerina
Marls and Luc aces ¸ ti Sandstone, Lower Menilite, Bituminous Marl,
Lower Dysodilic Shale with Kliwa Sandstones, Upper Dysodilic
Shale and Menilite and, finally, Gura S ¸ oimului formations (Micl aus ¸
et al., 2010). In our study area from the Cuejdiu basin, the Vrancea
Nappe crops out in the Bistrit ¸ a tectonic half-window (B ancil a, 1958;
Grasu et al., 1988) and the EarlyeLate Cretaceous deposits belong
to the S arata and Leps ¸a Formations (Guerrera et al., 2012).
The specimens of Orestella under review originate from the
* Corresponding author.
E-mail address: sacit.ozer@deu.edu.tr (S.
€
Ozer).
Contents lists available at ScienceDirect
Cretaceous Research
journal homepage: www.elsevier.com/locate/CretRes
http://dx.doi.org/10.1016/j.cretres.2014.09.003
0195-6671/© 2014 Elsevier Ltd. All rights reserved.
Cretaceous Research 52 (2015) 73e82