Abstract
The morphological basis of diabetic nephropathy has been
studied using light and electron microscopy. Kidneys of
streptozotocin-induced diabetic rats were examined on the
light microscope at 4 weeks and 8 months after induction of
diabetes mellitus. In addition, the 8-month diabetic kidneys
were examined with the electron microscope.
Renal hypertrophy was evidenced by the increase in the
weight of kidneys of diabetic rats. Whilst the diabetic kidneys
were approximately twice as large after 4 weeks they were
only 30% larger compared to age-matched controls after 8
months of induction of diabetes.
After 4 weeks, light microscopy revealed dilated tubules
within the cortex of the diabetic kidneys. Light microscopy
showed a significant amount of destruction of the distal
convoluted tubules while electron microscopy revealed a
spectrum of damage that included basement membrane
thickening, loss of podocytic foot processes, disruption of
tubular basal infoldings and their related mitochondria and
fibrosis of the tubules 8 months after induction of diabetes.
It is concluded that renal hypertrophy persists after a pro-
longed occurrence of diabetes but the extensive damage and
loss of renal tissue including the loss of the foot processes
of podocytes might be partly responsible for the clinical pre-
sentation of diabetic nephropathy.
Keywords: Diabetes, glomerulus, morphology, nephropathy,
rat kidney, streptozotocin.
Introduction
The kidney has become an important organ for the investi-
gation of the pathophysiology of diabetes mellitus and its
complications and several animal experiments have been
conducted to reproduce the typical human diabetic lesions of
nephromegaly, mesangial expansion and basement mem-
brane thickening. Of these animal models, the streptozotocin
(STZ) rat model has been most extensively studied to eluci-
date the morphological, physiological and biochemical
changes in the diabetic kidney.
In the STZ rat model, functional and structural features of
nephropathy appear within 6 months (Hirose et al., 1982) but
indications of renal disease such as an increase in glomeru-
lar filtration rate [GFR], renal plasma flow and renal hyper-
trophy develop within a week of induction of the disease
(Seyer-Hansen, 1983, 1987; Allen et al., 1990; Bach and
Jerums, 1990). The underlying mechanisms of the increase
in GFR (hyperfiltration), however, still remain unknown
inspite of intense research in this field. The association
between renal hypertrophy and hyperfiltration has also been
described in man (Mogensen & Andersen, 1973). It has been
shown that there is a progressive rise in rat urinary albumin
excretion (Cooper et al., 1988) analogous to the phases of
albuminuria (microalbuminaemia to macroalbuminaemia)
seen in man (Mogensen et al., 1983) and that while protein-
uria is evident in most diabetic animals, it is not as abundant
as in humans. It has been suggested that this major sieving
defect may be due to the loss of selective macromolecular
size permeability by the glomerular capillary (Michels et al.,
1982), possibly related to diminished heparan sulfate syn-
thesis (Rohrbach, 1986) resulting in reduced glomerular
basement membrane (GBM) barrier density and ionic charge
(Van den Born et al., 1995). Ultrastructural changes observed
in both diabetic animals and humans include thickening of
GBM and mesangial expansion (Steffes & Mauer, 1984;
Cooper et al., 1988). Several studies have also shown that the
synthesis of various mesangial and glomerular components
is considerably enhanced in diabetes (Fukui et al., 1993;
Accepted: 6 September, 2001
Address correspondence to: Prof. A. Adem, Dept. of Pharmacology, Faculty of Medicine and Health Sciences, P.O. Box 17666, Al Ain,
United Arab Emirates University, Tel.: 971-3-7039522; Fax: 971-3-7672033; E-mail: abdu.adem@uaeu.ac.ae
Morphological Changes in the Rat Kidney Following
Long-Term Diabetes
E.N. Obineche
1
, E. Mensah-Brown
2
, S.I. Chandranath
3
, I. Ahmed
2
, O. Naseer
3
and A. Adem
3
Departments of Internal Medicine
1
Anatomy
2
and Pharmacology
3
, Faculty of Medicine and Health Sciences, United Arab
Emirates University, Al Ain, United Arab Emirates
Archives of Physiology and Biochemistry 1381-3455/01/10903-241$16.00
2001, Vol. 109, No. 3, pp. 241–245 © Swets & Zeitlinger
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