ICANCERESEARCH 55. 1417—1422. AprilI, 19951
Advances in Brief
Immunohistochemical Quantitation of Polycyclic Aromatic Hydrocarbon-DNA
Adducts in Human Lymphocytes'
Grazyna Motykiewicz,Ewa Matusecka, Ewa Grzybowska, MieczyslawChorazy, Yu-Jing Zhang, Frederica P. Perera,
and Regina M. Santella2
Department of Tumor Biology. In.uitute of Oncology, 44-100 Gliwice, Poland (G. M., E. M., E. G., M. CI, and Cancer Center/Division of Environmental Sciences, Columbia
School of Public Health, New York, New York 10032 (1'-). 1, F. P. P., R. M. S.)
Refs. 2—5). They include 32P-postlabeling, gas chromatography/mass
spectrometry, fluorescence, and immunoassays (ELISA) using spe
cific antisera. Most methods are not applicable to small amounts of
blood or tissue, a major limitation in biomonitoring studies.
Antisera directed against PAH-DNA adducts can also be used for
the immunocytochemical localization of adducts in individual cells.
This approach has been successfully applied to mouse cells (6) and
tissues (7, 8) as well as human WBC treated in vitro with B[a]P (8).
Pilot immunofluorescence studies on bronchial cells from a smoker
have also been reported (9). Application of immunoperoxidase tech
niques have been applied to tissues and oral mucosa cells of smokers
and nonsmokers (10, 11).
Here we report on an immunofluorescence technique for visualiza
tion and quantitation of PAH-DNA adducts in individual human
lymphocytes. The formation of BPDE and related PAH-DNA adducts
was studied in lymphocytes obtained from coke-factory workers in
Silesia, Poland, residents from the vicinity of the cokery, and inhab
itants of a rural region in Poland with PAH levels estimated to be
10-fold lower than in Silesia. This study builds upon our earlier results
on aromatic adducts and PAH-DNA adducts measured on isolated
blood DNA by 32P-postlabeling and ELISA (12). Both methods
showed DNA adducts to be significantly elevated in the Silesian
groups (cokery workers and subjects living in the surrounding area)
compared to rural controls, with only a slight difference between
cokery workers and local controls. In addition to DNA adducts,
chromosomal aberrations and Sister chromatid exchanges were signif
icantly elevated in occupationally and environmentally exposed
residents of Silesia compared to controls (13, 14).
Materials and Methods
Chemicals. BPDE (444 mCi/mmol) was supplied by Chemsyn Science
Laboratories (Lenexa, KS). RNase, proteinase K, P1, Histopaque-1077, and
1,4-phenylenediamine were purchased from Sigma Chemical Co. (St. Louis,
MO).Biotinylatedgoatanti-rabbitlgGandfluorescein-conjugatedstreptavidin
was from Boehringer Mannheim (Indianapolis, IN). FCS was obtained from
HyCloneLaboratories, Inc. (Logan,UT),and DMEMmediumwaspurchased
from GIBCO-BRL Life Technologies (Gaithersburg, MD). Tissue culture
chambers were supplied by NUNC (Naperville, IL).
Subjects and Blood Samples. Subjects included 16 healthy males from
each exposure group as well as 16 rural controls. Donors were selected on the
basis of their occupation; environmental and control groups did not include
cokery workers or any other workers connected with the processing of coal.
Additional information on cigarette smoking, diet, X-ray exposure, current
medication, and family history of cancer was also collected. Samples were
taken in September 1992 and 1993, which is considered the summer season.
Therefore, the exposure was not influenced by the additional impact of PAH
from domestic heating in the winter.
Treatment of Cells with BPDE. The immunostaining method was estab
lished, and the measurements were calibrated on cultured C3H lOT½cells
treated with BPDE. Cells were plated in eight-chambered slides and main
tamed in DMEM medium supplemented with 10% FCS. After 24 h, cells were
1417
Abstract
The formation of polycydlic aromatic hydrocarbon-DNA adducts was
studied in @pheral blood lymphocytes obtained from men with occupa
fional and environmental exposure. Subjects Included coke factory workers,
residents from the vicinity of the cokery, and niral region inhabitants (16
individuals in each exposure group). Adducts were determined by immuno
histochemical analysis using a polyclonal antiserum recognizing benzo
(a)pyrene and related polycydicaromatic hydrocarbon dial epoxide-DNA
adducts, a biotinylated secondary antiserum, and @ptavidin-conjupted
F1TC. Propidium iodide was used to quantitate nuclear DNA. Dual fluores
cence intensifies were simultaneously measured with a Zeiss Axiovert micro
scope and a Blo-Rad MRC-600 argon laser scanning confocal attachment.
Adducts were significantly elevated (P < 0.001) in both occupational and
envlromnental groups, as compared to the rural control group by Mann
Whitney U test. The distribution of the data indicated the existence of cells
with relatively higher adduct Ievel& The percentages ofthese so called â€oehigh
er adduct-level cells― were 13.6, 11.5, and 3.7 in cokery workers, environ
mentally exposed Individuals, and rural controls, respectively. The immuno
histochemical method allows visualization and relative quantitation of
polycydlicaromatic hydrocarbon-DNA adducts In Individual lymphocytes. It
requires a much smaller amount of blood than the previously used @
@bellngand ELISA methods, which used isolated bulk DNA. It can also
be used for adduct quantitadon In biopsy materiaL The results of this pilot
study indicate that this technique is a promising addition to biomonitoring
sftidIe@
Introduction
PAH3 are common environmental pollutants, and their uptake has
been causally associated with the induction of cancer. As a result of
coal burning and/or processing, PAH are among the most prevalent
mutagenic air pollutants in Silesia, Poland (1). The highest levels are
found in the occupational environment, i.e., coke-oven plants
(0.12—89.58 @ of B[a}P/m3 of air). Chronic exposure to lower levels
of PAH (1.0—55.3ng B[a]P/m3) occurs in the general environment of
Silesia and mainly originates from coal-based heavy industry, power
plants, and individual domestic heating, as well as automobile traffic.
Among the known molecular effects of PAH is the ability to bind
covalently to DNA, giving rise to PAH-DNA adducts. A number of
methods have been developed for quantitation of adducts, resulting
from occupational or environmental exposure to PAH (reviewed in
Received 1/18/95; accepted 2/20/95.
Thecostsof publicationof thisarticleweredefrayedin partby thepaymentof page
charges. This article must therefore be hereby marked advertisement in accordance with
18 U.S.C. Section 1734 solely to indicate this fact.
I This work was supported by Grants CA21 1 1 1 and ESO5249 from the NIH, Grant 6
P20706405p01 from the Committee for Scientific Research (Komitet Badan Navkowych)
to M. C., and a fellowship to G. M. from the National Cancer Institute Short-Term
Scientist Exchange Program.
2 To whom requests for reprints should be addressed, at Columbia University, 701
West 168th St., New York, NY 10032.
3 The abbreviations used are: PAH, polycycic aromatic hydrocarbons; B[aJP, benzo
(a)pyrene;BPDE,(±)7,8-dihydroxy-9,lO-epoxy-7,8,9, lO-tetrahydrobenzo(a)pyrene (the
anti isomer); P1, propidium iodide.
Research.
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