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106 Mini-Reviews in Medicinal Chemistry, 2013, 13, 106-123
Snake Venom Induced Local Toxicities: Plant Secondary Metabolites as an
Auxiliary Therapy
M. Sebastin Santhosh, M. Hemshekhar, K. Sunitha, R.M. Thushara, S. Jnaneshwari,
K. Kemparaju and K.S. Girish
*
Department of Studies in Biochemistry, University of Mysore, Manasagangothri, Mysore-570 006, Karnataka, India
Abstract: Snakebite is a serious medical and socio-economic problem affecting the rural and agricultural laborers of
tropical and sub-tropical region across the world leading to high morbidity and mortality. In most of the snakebite incidences,
victims usually end up with permanent tissue damage and sequelae with high socioeconomic and psychological impacts.
Although, mortality has been reduced markedly due to anti-venom regimen, it is associated with several limitations. Snake
venom metalloprotease, hyaluronidase and myotoxic phospholipase A
2
are the kingpins of tissue necrosis and
extracellular matrix degradation. Thus, inhibition of these enzymes is considered to be the rate limiting step in the
management of snakebite. Unfortunately, tissue necrosis and extracellular matrix degradation persists even after the
administration of anti-venom. At present, inhibitors from snake serum and plasma, several synthetic compounds and their
analogs have been demonstrated to possess anti-snake venom activities, but the use of plant metabolites for this purpose
has an added advantage of traditional knowledge and will make the treatment cheaper and more accessible to the affected
population. Therefore, the clinical and research forums are highly oriented towards plant metabolites and interestingly,
certain phytochemicals are implicated as the antibody elicitors against venom toxicity that can be exploited in designing
effective anti-venoms. Based on these facts, we have made an effort to enlist plant based secondary metabolites with anti-
ophidian abilities and their mechanism of action against locally acting enzymes/toxins in particular. The review also
describes their functional groups responsible for therapeutic beneficial and certainly oblige in designing potent inhibitors
against venom toxins.
Keywords: Hyaluronidase, Local toxicity, Phytochemicals, PLA
2,
Snake venom, SVMPs, Synthetic drugs.
INTRODUCTION
Snake envenomation is a serious medical and socio-
economic crisis affecting the healthy individuals like
children, rural population, specially the agricultural laborers
in tropical and sub-tropical regions. Snakebite has become a
nightmare for the inhabitants of southern Asia, northern
Africa, Latin America and the Middle East nations. The
affected regions experience high mortality and morbidity
rates because of poor access to health services. In addition,
most of the snakebite victims end up with permanent tissue
damage and sequelae, with high socioeconomic and
psychological impacts. Although the accurate statistics of
snakebite incidence is difficult to estimate, available data
reported of about 5 million cases of snakebites globally per
annum, of which 3.75 million people severely get affected
affected and results in 1.25 million deaths [1, 2]. About
2,50,000 snake bite incidences are recorded in India every
year and resulting in more than a 100,000 deaths and most of
them go unnoticed [3, 4]. In view of this, snakebites are
being considered and recognized as a major public health
hazard across the world and may therefore be appropriately
categorized as a neglected tropical disease by world health
organization [1].
*Address correspondence to this author at the DOS in Biochemistry,
University of Mysore, Manasagangothri, Mysore-570 006, Karnataka, India;
Tel: +91-9972268633; E-mail: ksgbaboo@yahoo.com
Of the 3000 known species of snakes 410 are venomous
and are particular to tropical and subtropical regions of the
world. There are about 53 venomous snakes in India of
which majority of the bites and mortality are attributed to
species like Naja naja (spectacled cobra), Daboia russelli
(Russell's viper), Bungarus caeruleus (common krait), Echis
carinatus (saw-scaled viper), and Ophiophagus hannah
(king cobra). The composition of venom varies with the
species and the variation was observed due to the differences
in age, sex, habitat, diet and season. Hence, the individual
venoms would exert varied spectrum of pharmacological
properties [5, 6].
Snake venom is an awfully composite concoction of
pharmacologically active enzymatic and non-enzymatic
protein and peptide toxins that assist in prey immobilization
mainly as hypotensive, paralytic and digestive aids. There
are more than twenty different enzymes present in venom
including phospholipases, snake venom metalloproteases
(SVMPs), snake venom hyaluronidases (SVHYs), serine
proteases, phosphatases (acid and alkaline), acetyl
cholinesterase, transaminase, phosphodiesterase, nucleotidase,
ATPase and nucleosidases [7].
Snake envenomation occurs due to subcutaneous/
intramuscular injection of venom into the human victims
resulting in complicated pharmacological effects that depend
on the combined and synergistic action of toxic and non-toxic
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