890 M. SARI ET AL.
Copyright © 2006 John Wiley & Sons, Ltd. Flavour Fragr. J. 2006; 21: 890–898
DOI: 10.1002/ffj
FLAVOUR AND FRAGRANCE JOURNAL
Flavour Fragr. J. 2006; 21: 890–898
Published online 29 June 2006 in Wiley InterScience
(www.interscience.wiley.com) DOI: 10.1002/ffj.1738
Chemical composition, antimicrobial and antioxidant
activities of the essential oil of several populations
of Algerian Origanum glandulosum Desf.
Madani Sari,
1
Daniela M. Biondi,
2
Mohamed Kaâbeche,
3
Giuseppina Mandalari,
4
Manuela D’Arrigo,
4
Giuseppe Bisignano,
4
Antonella Saija,
4
Carmelo Daquino
2
and Giuseppe Ruberto
2
*
1
Département de Biologie, Faculté des Sciences et Sciences de l’Ingénieur, Université Mohamed Boudiaf, 28000 M’Sila, Algérie
2
Istituto del C.N.R. di Chimica Biomolecolare, Via del Santuario, 110 I-95028 Valverde CT, Italy
3
Département de Biologie, Faculté des Sciences, Laboratoire d’Ecologie Végétale, Université Ferhat Abbas, 19000 Setif, Algérie
4
Dipartimento Farmaco-Biologico, Università di Messina, Contrada Annunziata, I-98128 Messina, Italy
Received 14 June 2005; Revised 12 November 2005; Accepted 30 January 2006
ABSTRACT: Essential oils extracted by hydrodistillation from the aerial parts of 23 samples of Algerian Origanum
glandulosum Desf. were analysed by gas chromatography (GC) and GC–mass spectrometry (MS). Overall, 30 components
have been fully characterized. However, all the oils were characterized by the predominance of four components, thymol
(18.5–73.1%), carvacrol (7.6–72.6%), p -cymene (1.7–18.5%) and
γ
γ
γ -terpinene (1.1–18.7%). Cluster analysis of the iden-
tified components allowed us to establish the presence of three main groups, characterized by carvacrol, thymol and com-
parable amounts of both compounds, respectively. The free radical scavenging activity of essential oils was determined
by the 1,1-diphenyl-2-picrylhydrazyl (DPPH) model system. The SC
50
(scavenging concentration) values were in the range
16.2–26.7 μg/ml, representing a good antioxidant effectiveness. The roles of thymol and carvacrol, the main components
of all oils, were estimated by measuring their stoichiometric factors. The essential oils were also evaluated for their
antimicrobial activity by the agar disc diffusion method and the determination of minimum inhibitory concentration (MIC)
against six standard strains (Escherichia coli, Pseudomonas aeruginosa, Staphylococcus aureus, Enterococcus hirae, Candida
albicans, Candida tropicalis). All microbial strains employed (Gram-positive and Gram-negative bacteria and yeasts)
showed a fairly similar degree of susceptibility to the essential oils under investigation, although no evident difference
was observed in their sensitivity. Furthermore, a similar level of toxicity was observed for all oils examined, with MIC
values of 31.25–125.00 μg/ml. Finally, the addition of the emulsifier Tween 80 to the oil or to the agar markedly decreases
the antimicrobial activity of the essential oils against all microbial strains employed, thus suggesting that the antimicro-
bial activity of the essential oils is dependent on the physicochemical characteristics of their components and also on the
microbial strains employed. Copyright © 2006 John Wiley & Sons, Ltd.
KEY WORDS: Origanum glandulosum Desf.; Lamiaceae; antimicrobial and antioxidant activities; essential oil; cluster analysis;
thymol; carvacrol
* Correspondence to: G. Ruberto, Instituto del C.N.R. di Chimica
Biomolecolare, Via del Santuario, 110 I-95028 Valverde CT, Italy.
E-mail: giuseppe.ruberto@icb.cnr.it
Contract/grant sponsor: CNR, Rome, Italy.
Introduction
The genus Origanum (family Lamiaceae) comprises
about 38 species, most of which are indigenous to the
Mediterranean, Euro-Siberian and Irano-Siberian regions,
although 75% of them are restricted to the eastern
Mediterranean area.
1
Due to the variability in chemical
and aroma composition, Origanum plants are widely used
as a culinary herb, to flavour food products and alcoholic
beverages
2
and for their pharmacological properties,
including antibacterial, antioxidant, antithrombin and
antihyperglycaemic activities.
3–6
Origanum species have continuously been studied
for their essential oil composition as potential sources
of antimicrobial and antioxidant natural agents. On the
basis of these studies, as well as the prevalence of the
main compounds, Origanum taxa can be divided in three
groups: (a) linalool, terpinen-4-ol and sabinene hydrate
group; (b) carvacrol and/or thymol group; (c) sesquiter-
penes group.
7
As previously demonstrated, the content of
essential oil and extracts of Origanum species, like other
medicinal plants with antimicrobial, antioxidant and other
biological activities, may change depending on the differ-
ences in cultivation, origin, vegetative stage and growing
season of the plants.
8,9
Origanum glandulosum Desf. (Lamiaceae) is an
endemic herbaceous or sub-ligneous plant in the Algerian
and Tunisian areas,
10
being one of the most widespread
plants of the Setif region of north-eastern Algeria,
11