A review of the upper Campanian vertebrate site of Armu ~ na (Segovia Province, Spain) A. P  erez-García a, b, * , F. Ortega a , A. Bolet c , F. Escaso a , A. Houssaye d , J. Martínez-Salanova, C. de Miguel Chaves a , P. Mocho a, e , I. Narv  aez a , M. Segura f , A. Torices g , D. Vidal e , J.L. Sanz e a Grupo de Biología Evolutiva, Facultad de Ciencias, UNED, Paseo Senda del Rey, 9, 28040 Madrid, Spain b Centro de Geologia, Faculdade de Ci^ encias da Universidade de Lisboa (FCUL), Edificio C6, Campo Grande, 1749-016 Lisbon, Portugal c Institut Catal a de Paleontologia Miquel Crusafont, Escola Industrial 23, E-08201 Sabadell, Barcelona, Spain d UMR 7179 CNRS/Mus eum National d'Histoire Naturelle, D epartement Ecologie et Gestion de la Biodiversit e, 57 rue Cuvier CP-55, 75000 Paris, France e Unidad de Paleontología, Departamento de Biología, Facultad de Ciencias, Universidad Aut onoma de Madrid, 28049 Madrid, Spain f Grupo de Investigaci on IberCreta, Facultad de Ciencias, Universidad de Alcal a de Henares, 28871 Alcal a de Henares, Madrid, Spain g Department of Biological Sciences, Faculty of Science, CW405 Biological Sciences Building, University of Alberta, Edmonton, Alberta, Canada article info Article history: Received 28 April 2015 Received in revised form 1 August 2015 Accepted in revised form 16 August 2015 Available online xxx Keywords: Late Cretaceous Iberian Peninsula Armu~ na Vertebrates Reptiles abstract The Upper Cretaceous outcrops of Armu~ na (Segovia Province, Spain) yielded relatively abundant material of vertebrates during prospection and excavation in the second half of the 1980s. However, little has been published on these remains. A new analysis of the specimens from this upper Campanian site reveals the presence of some clades in the site for the first time (e.g., Dortokidae, Anguimorpha, Mosasauroidea). Furthermore, the material of the clades previously recognized there has been reviewed and described in more detail, with some previous systematic attributions confirmed and others refuted. Consequently, a relatively high local diversity has been identified. New taxa (i.e., a member of Anguimorpha and a eusuchian crocodyliform) are identified in Armu~ na, coexisting with other taxa previously described in other sites from the Iberoarmorican Realm. The vertebrates from Armu~ na confirm that the fauna from the Upper Cretaceous of the Iberian Peninsula is composed of a mixture of European endemic clades and lineages shared with other continents such as North America (e.g., anguimorphs) and Africa (e.g., bothremydids). © 2015 Elsevier Ltd. All rights reserved. 1. Introduction Excavations at several European vertebrate sites for the last three decades and subsequent studies on many specimens found have allowed characterizing the Late Cretaceous European faunas as some of the best known in the global record from this age (Csiki- Sava, Buffetaut, } Osi, Pereda-Suberbiola, & Brusatte, 2015). Recently, summaries on the faunal composition of some of the most repre- sentative Iberian Upper Cretaceous sites, such as the Spanish sites of Lo Hueco (Cuenca) and La~ no (Condado de Trevi ~ no), have just been published (see Ortega et al., 2015 and Pereda-Suberbiola et al., 2015). The Upper Cretaceous site of Armu~ na (upper Campanian of the Vegas de Matute Formation, Central System, Segovia Province) was discovered and excavated in the second half of the 1980s. Despite having yielded abundant and diverse vertebrates (Buscalioni and Martínez-Salanova, 1990), only a few of the remains from this site have been thoroughly studied. The only published faunal list on the vertebrates from Armu~ na was preliminar, and was published more than two decades ago in an abstracts volume (see Buscalioni and Martínez-Salanova, 1990). The scarce available in- formation on the taxa identified in Armu~ na needed to be reviewed taking into account the current knowledge. In addition, several clades represented there remained unpublished until now. The aim of this paper is to characterize the vertebrate fauna from Armu~ na by thoroughly describing and analyzing the fossils. The systematic of the vertebrates recorded there has been analyzed, as well as their biostratigraphic and biogeographic con- texts. In addition, the presence of vertebrate remains in Carbonero el Mayor, another upper Campanian locality of the Segovia Province * Corresponding author. Grupo de Biología Evolutiva, Facultad de Ciencias, UNED, Paseo Senda del Rey, 9, 28040 Madrid, Spain. E-mail address: paleontologo@gmail.com (A. P erez-García). Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes http://dx.doi.org/10.1016/j.cretres.2015.08.008 0195-6671/© 2015 Elsevier Ltd. All rights reserved. Cretaceous Research xxx (2015) 1e33 Please cite this article in press as: P erez-García, A., et al., A review of the upper Campanian vertebrate site of Armu~ na (Segovia Province, Spain), Cretaceous Research (2015), http://dx.doi.org/10.1016/j.cretres.2015.08.008