Neurons of the human dentate nucleus: box-count
method in the quantitative analysis of cell
morphology
Dušica L. Mari
Department of Anatomy
Medical faculty, University of Novi Sad, Serbia
E-mail: maricduska@gmail.com
Nebojša T. Miloševi
Department of Biophysics
Medical faculty, University of Belgrade, Serbia
E-mail: mtn@med.bg.ac.rs
Herbert F. Jelinek
Department of Biomedical Engineering, Khalifa University
of Science, Technology and Research, UAE
E-mail: hjelinek@csu.edu.au
Katarina Rajkovi
Laboratory for Image Analysis in Medicine
Medical faculty, University of Belgrade, Serbia
E-mail: katar1970@gmail.com
Abstract-The morphology of neurons from the human
dentate nucleus was analyzed estimating the size and shape
of the dendritic field, shape of the neuron, space-filling
property and the degree of dendrite aberrations. Among
them, the last three morphological properties were
investigated using the most popular technique of fractal
analysis: the box-count method. The box dimensions of
binary images and dendritic field area were statistically
investigated in order to test whether the binary box
dimension can quantify the size of the neuron. The same
analysis was carried out using the box dimension of outline
images and image circularity. The parameters, presented
in this study have proved to be a useful means for
quantifying the morphology of dentate neurons as they
provide a robust means of differentiating between neuron
subtypes in the dentate nucleus. The findings of the
present study are in accordance with previous qualitative
data.
Keywords-Box dimension, Cerebellar cortex, Fractal
analysis, Neuronal Morphology, Two dimensional images
I. INTRODUCTION
The dentate nuclei, the largest and phylogenetically most
recent of the cerebellar nuclei are located in the right and left
hemisphere of the cerebellar white matter and play a role as a
major relay center between the cortex and other parts of the
brain. The dentate nuclei are responsible for the planning,
initiation and control of volitional movements [1].
Although the geometry of the neuronal soma and dendritic
fields has been previously studied in the dentate nuclei, a
quantitative classification of neuronal types and their analysis
was not satisfactorily explored. According to various
qualitative studies [2-5], the dentate nuclei consist of large
neurons with long axons and small neurons with short axons.
In the above cited publications, the neuronal morphology and
determination of neuron types were primarily based on
qualitative descriptions of the shape and size of the cell bodies
[6].
One of the earliest descriptions of neurons in the human
dentate nuclei was based on Golgi stained neurons of the
neonatal human cerebellum [2,3]. Recent descriptions offer
some new data concerning the neuronal morphology of the
dentate nuclei at various gestational ages [7-12] and early
postnatal ages [8,13].
In adult human dentate nuclei, some authors described the
presence of two cell types [5], while other have proven that the
class of large neurons could be further subdivided into four
types: central, border, asymmetrical and fusiform multipolar
neurons [14,15].
Thus, we undertook the present study with the aim to
investigate features of the large neurons in prenatal and adult
human dentate nuclei. For that purpose, we estimated the size
and shape of the dendritic field, shape of the neuron, space-
filling property and the degree of dendrite aberrations of the
neurons.
The shape of the neuron, space-filling property and the
degree of dendrite aberrations were estimated using the box-
count method of fractal analysis. In addition, box dimensions
of binary images and dendritic field area, as well as the box
dimension of outline images and image circularity, were
statistically investigated in order to test whether box
dimensions can quantify the size of the neuron and the shape
of the dendritic field.
II. MATERIALS AND METHODS
Camera lucida drawings of forty-two Golgi impregnated
neurons were taken from dentate nuclei of human fetuses
ranging from 24 to 41 gestational weeks (gw). The fetal
cadavers were collected either from hysterectomy or autopsy
[11]. The gestational age of the fetuses was estimated by
taking into consideration the fetal crown rump length,
biparietal diameter, the foot length as well as the maternal
history of last menstrual period and the size of the uterus [16].
2013 19th International Conference on Control Systems and Computer Science
978-0-7695-4980-4/13 $26.00 © 2013 IEEE
DOI 10.1109/CSCS.2013.33
319
2013 19th International Conference on Control Systems and Computer Science
978-0-7695-4980-4/13 $26.00 © 2013 IEEE
DOI 10.1109/CSCS.2013.33
319