Journal of Applied Microbiology 1997, 82, 729–734
Determination of bacteria associated with reared turbot
(Scophthalmus maximus) larvae
A.R. Blanch, M. Alsina, M. Simo ´ n and J. Jofre
Department of Microbiology, University of Barcelona, Barcelona, Catalonia, Spain
5796/06/96: received 10 June 1996, revised 25 October 1996 and accepted 28 October 1996
A.R. BLANCH, M. ALSINA, M. SIMO
´
N AND J. JOFRE. 1997. In order to extend our knowledge
of the presence of bacteria in hatcheries and their influence on rearing
performance, the aerobic and facultative bacterial flora associated with farmed turbot
larvae were studied in relation to the microflora of the water and diets. A settlement
of specific groups of bacterial populations was found in the gut of the larvae. A clear
succession of bacterial phenotypes was also observed from day 1 to day 90 post-hatching.
Oxidative Gram-negative rods were predominant at the initial stages, whereas some
phenotypes of Vibrio were frequent at the final stages. A high heterogeneity of Vibrio species
was observed in the intermediate period when the highest mortalities of turbot larvae
occur.
INTRODUCTION 1977; Tanasomwang and Muroga 1988). However, others
report that the bacterial flora of farmed fish was not consistent
The source, the characteristics and the biological significance
with that of water or the fish diet (Campbell and Buswell
of the microbial flora in the intestine of marine fish are not
1983; Muroga et al. 1987; Nicolas et al. 1989). Bacteria of the
well known. An understanding of the features and the role of
genus Vibrio seem to constitute the dominant flora of the
the indigenous microflora of marine fish larvae may help to
intestine of marine fish (Sakata et al. 1980; Onarheim and
improve feeding and other conditions for the intensive rearing
Raa 1990), independently of the bacterial content in the water
of fish. Several functions have been assigned to the resident
or in the food ingested (Campbell and Buswell 1983; Muroga
gut microflora. These include positive effects on nutrition
et al. 1987; Nicolas et al. 1989). Quantitative studies have been
(Fong and Mann 1980; MacDonald et al. 1986; Rimmer
performed to evaluate the influence of associated bacteria on
and Wiebe 1987) and possible prevention of colonization by
the survival of turbot larvae (Nicolas et al. 1989; Westerdahl
bacterial pathogens (Onarheim and Raa 1990; Westerdahl et
et al. 1991). However, the diversity of the bacterial population
al. 1994; Bergh 1995). The latter may involve inactivation
in water, food and larvae at different stages of rearing has not
of pathogenic bacterial toxins or metabolites, production of
been considered. We present a study performed in a turbot
bacteriocins or inhibitory factors, changes in pH and redox
hatchery, in which aerobic and facultative bacteria have been
potential, competition with pathogens for nutrients and
isolated and identified from these different origins. The
adhesion sites and stimulation of non-specific immunity (Jong
results may facilitate the prediction of some of the charac-
1993). Aquaculture of marine fish has resulted in intensive
teristics of the microbiota that most frequently colonize the
systems for egg incubation and hatching. In such systems,
intestinal tract of turbot larvae in intensive rearing.
newly hatched larvae, which for a short time have a sterile
intestinal tract (Wood 1967), are exposed to an environment,
with variable bacterial populations (Hansen and Olafsen
MATERIALS AND METHODS
1989). Moreover, such bacterial populations may be different
to those of the natural environment. Some authors report
Fish larvae
that the intestinal microflora of fish reflects the bacterial
Turbot larvae were sampled at Tinamenor Hatchery (San-
content of ingested food and of the water (Seki 1969; Horsley
tander, Spain) from day 1 after eclosion of eggs to day 100,
when they were ready to leave the hatchery. Samples were
Correspondence to: Dr Anicet R. Blanch, Department of Microbiology,
taken from the production lots kept under the usual rearing
University of Barcelona, Diagonal 645, 08028 Barcelona, Catalonia, Spain (e-
mail: anicet@porthos.bio.ub.es). conditions for this hatchery. Six thousand-litre tanks main-
© 1997 The Society for Applied Bacteriology