PART V, REVISED, CHAPTER 12: GLOSSARY OF THE HEMICHORDATA JÖRG MALETZ, 1 DENIS E. B. BATES, 2 EDSEL D. BRUSSA, 3 ROGER A. COOPER, 4 ALFRED C. LENZ, 5 JOHN F. RIVA, 6 BLANCA A. TORO, 7 and YUANDONG ZHANG 8 [ 1 Institut für Geologische Wissenschaften, Berlin, Germany, Yorge@zedat.fu-berlin.de; 2 Institute of Geography and Earth Sciences, Aberystwyth University, UK, deb@aber.ac.uk; 3 Deceased; 4 DSIR GEO, New Zeandand Geological Survey, R.Cooper@gnx.cri.nz; 5 University of Western Ontario, Canada, aclenz@uwo.ca; 6 INRS-ETA, Québec, Canaba, john_riva@ete.enrs.ca; 7 CICTERRA, Ciudad Universitaria, Cordoba, Argentina, btorogr@ mendoza-conicet.gob.ar; 8 Nanjing Institute of Geology and Palaeontology; Academica Sinica, Nanjing, China, ydzhang@nigpas.ac.cn] PREFACE The terminology of the Hemichordata used herein differs considerably from the terminology used in the previous versions of the “Graptolite Treatise” (BULMAN, 1955, 1970), which were focused entirely on the fossil members of the Graptolithina and the few known tube-bearing extant and extinct Pterobranchia. Both classes were regarded as independent groups of the Hemichordata, but they are now included as a single taxon, the Pterobranchia, with the Graptolithina and the Cephalodiscida as two subclades (MITCHELL & others, 2013). A number of terms had to be changed due to a better understanding of the construction of the graptolite tubarium and the phylogenetic relationships within the Pterobranchia, the main group of fossilized hemichordates, in an attempt to homogenize the terminology of these previously separated groups. The term tubarium, for example, is rein- troduced to the pterobranch terminology to describe the organic housing or domi- cile of all Pterobranchia, instead of using the term rhabdosome for the fossil Ptero- branchia (Graptolithina in BULMAN, 1955, 1970) and the coenecium for the extant members (Pterobranchia sensu BULMAN, 1955, 1970). LANKESTER (1884) initially introduced the term tubarium to describe the housing of the extant Pterobranchia, as he considered the terms coenecium or zooe- cium inappropriate since they described the housing construction of bryozoans, to which the Pterobranchia were referred initially. Tubarium describes the housing construc- tion of extant and extinct Pterobranchia more independently; it also describes it more precisely as being formed by glandular secre- tions and can easily be used for all members. It is therefore extended herein to include the homologous glandular constructions of the domiciles of fossil graptolite taxa. Other terms, such as the periderm, have been eliminated due to constructional considerations. The term periderm, intro- duced by WIMAN (1895), suggests a dermal construction of the tubarium, which is now known to be incorrect. The tubarium is formed from glands on the head shield of the zooids and is not a dermal construction as is the coenecium of a bryozoan. The tubarium can perhaps be compared with the formation of a hornet’s nest, though a hornet’s nest is largely constructed with foreign material. A comparison with hydrozoan colonies also falls short, as these are often covered by an organic exoskeleton, the perisarc (also iden- tified as the periderm in HYMAN, 1940, p. 400), which is secreted by the epidermis of the advancing stolon (e.g., BERKING, 2006). Early graptolite literature is written in a number of languages, including Chinese, Czech, English, French, German, Latin, Norwegian, Spanish, and Swedish. Thus, paleontological terms describing grapto- lite features were created in a variety of languages. Many terms originally proposed in other languages were translated into English before they became the standard, © 2014, The University of Kansas, Paleontological Institute, ISSN 2153-4012 Jörg Maletz, Denis E. B. Bates, Edsel D. Brussa, Roger A. Cooper, Alfred C. Lenz, John F. Riva, Blanca A. Toro, & Yuandong Zhang. 2014. Part V, Revised, Chapter 12: Glossary of the Hemichordata. Treatise Online 62:1–23, 6 fig.