Vaccine 33S (2015) B40–B43
Contents lists available at ScienceDirect
Vaccine
j o ur na l ho me page: www.elsevier.com/locate/vaccine
Review
Gaps in knowledge and prospects for research of adjuvanted vaccines
Robert Seder
a
, Steven G. Reed
b,*
, Derek O’Hagan
c
, Padma Malyala
c
, Ugo D’Oro
d
,
Donatello Laera
d
, Sergio Abrignani
e
, Vincenzo Cerundolo
f
, Lawrence Steinman
g
,
Sylvie Bertholet
d
a
Cellular Immunology Section, Vaccine Research Center, National Institute of Allergy and Infectious Diseases, Bethesda, MD, USA
b
Infectious Disease Research Institute, Seattle, WA, USA
c
Novartis Vaccines, Cambridge, MA, USA
d
Novartis Vaccines, Via Fiorentina 1, Siena, Italy
e
Instituto Nazionale Genetica Molecolare, Milano, Italy
f
Oxford University, Oxford, England, United Kingdom
g
Stanford University, School of Medicine, Palo Alto, CA, USA
a r t i c l e i n f o
Keywords:
Vaccine
Adjuvant
a b s t r a c t
A panel of researchers working in different areas of adjuvanted vaccines deliberated over the topic,
“Gaps in knowledge and prospects for research of adjuvanted vaccines” at, “Enhancing Vaccine Immunity
and Value” conference held in July 2014. Several vaccine challenges and applications for new adjuvant
technologies were discussed.
© 2015 Elsevier Ltd. All rights reserved.
1. Introduction
From a historical perspective, vaccines have been instrumental
in eradicating or drastically reducing the incidence of numerous
dreaded infectious diseases such as small pox, polio, tetanus,
diphtheria, cholera, rabies and typhoid [1]. Vaccines to combat
two major deadly infections, human papillomavirus [2] and
meningococcal disease [3], were also introduced in the recent
past, and ongoing research aims to eliminate many more diseases.
The need for optimizing adjuvant formulations for new vaccines
presents a number of challenges, though for most of these paths
forward are becoming evident. This workshop addressed issues
and potential solutions focusing on developing vaccines for HIV,
malaria, tuberculosis (TB), and cancer. Although certain challenges
are specific for each of these vaccine targets, some basic issues
hold constant. These include adjuvant access, the importance of
formulation, and the need for adjuvants that promote appropriate
T cell responses, whether as effectors or helpers of effective and
durable antibody responses.
A radical change has taken place in the evolution of vaccines
since Edward Jenner’s inoculation of cowpox matter as a small
pox vaccine in the 1800s in an eight-year-old boy [4]. This seminal
example using an attenuated viral vaccine to elicit long lived
and broad based humoral and cellular immunity highlights the
*
Corresponding author. Tel.: +1 206 381 0883.
E-mail addresses: sreed@idri.org, steve.reed@idri.org (S.G. Reed).
efficiency by which antigens and innate immunity induced by the
virus delivered synchronously is so effective. Since this discovery
many vaccine approaches have become more selective where
the science has emerged to encompass antigens in the form of
conjugates, toxoids and recombinant vectors, inactivated and live
attenuated vaccines and subunit vaccines [5].
In terms of proteins or inactivated viral productions, adjuvants
have been used to improve the efficiency, potency and durability
of immune responses [6]. There is extensive history of the use of
vaccine adjuvants, mostly inadvertently through the inclusion of
natural products in the form of live or inactivated virus or microbial
products, but more recently through intentional efforts to develop
vaccine components to enhance immune responses. Two such com-
ponents in wide use are aluminum salts, commonly referred to as
alum, and oil in water emulsions such as MF59 [7]. In addition to the
empirical first generation adjuvants such as Alum, MF59 and AS03
emulsions, second generation adjuvants are typically agonists of
toll like receptors (TLRs) and act by triggering signal transduction
pathways that lead to activation of innate and adaptive immune
cells. Though widely used as standalone adjuvants, both alum and
emulsions may be used with TLR (and other) ligands as important
formulation components. The most advanced example of this is the
TLR4 ligand MPL
®
which has been combined with alum in the GSK
Cervarix
®
vaccine [8].
MPL
®
is purified from endotoxin (lipopolysaccharide, LPS)
derived from Gram-negative bacteria [9]. Endotoxin has been
known as a potent stimulus of antibody responses, and exten-
sive biochemical, biophysical, and immunological studies were
http://dx.doi.org/10.1016/j.vaccine.2015.03.057
0264-410X/© 2015 Elsevier Ltd. All rights reserved.