MUTATION IN BRIEF
HUMAN MUTATION Mutation in Brief #737 (2004) Online
Carbonic Anhydrase II Deficiency Syndrome
(Osteopetrosis with Renal Tubular Acidosis and
Brain Calcification): Novel Mutations in CA2
Identified by Direct Sequencing Expand the
Opportunity for Genotype-Phenotype Correlation
Gul N. Shah
1
, Giuseppe Bonapace
1
, Peiyi Y. Hu
1
, Pietro Strisciuglio
2
, and William S. Sly
1
*
1
Department of Biochemistry, Saint Louis University School of Medicine, St. Louis, MO 63104;
2
Department of
Pediatrics, University of Catanzaro, 88100 Catanzaro, Italy
*Correspondence to: William S. Sly, M.D., Department of Biochemistry, Saint Louis University School of
Medicine, 1402 S. Grand Blvd., St. Louis, MO 63104; Tel.: 314-977-9201; Fax: 314-977-1183; E-mail:
slyws@slu.edu
Grant sponsor: National Institutes of Health (W.S.S.); Grant numbers: GM34182, DK40163, GM53405; Grant
sponsors: M.U.R. and C.O.F.I.N. (P.S.); Grant numbers: 980618306-006; MM06182533-006
Communicated by Elizabeth Neufeld
The carbonic anhydrase II (CA II) deficiency syndrome is an autosomal recessive disorder
that produces osteopetrosis, renal tubular acidosis, and cerebral calcification. Other features
include developmental delay, short stature, cognitive defects, and a history of multiple
fractures by adolescence. With one exception, all patients with osteopetrosis and renal
tubular acidosis examined have proven to have CA II deficiency. All CA II-deficient patients
analyzed have been found to have mutations in the CA2 gene. Previously, we used single
strand conformational (SSCP) analysis to identify exons to be sequenced from CA II-
deficient patients. In this report, we amplified all seven exons by PCR from genomic DNA
and directly sequenced the amplified products. Application of this method allowed
identification of eleven new mutations in 21 patients referred for confirmation of the
diagnosis of CA II deficiency. These mutations were scattered over the genome from exon 2
to 7. In two opportunities for prenatal diagnosis, one from cultured amniocytes and one
from chorionic villus biopsy, we demonstrated the general utility of the direct sequencing
method for prenatal DNA diagnosis. These studies expand our knowledge of the
heterogeneity in mutations underlying the CA II deficiency syndrome. © 2004 Wiley-Liss, Inc.
KEY WORDS: osteopetrosis; renal tubular acidosis; carbonic anhydrase II; CA2; prenatal diagnosis; cerebral calcification
INTRODUCTION
There are 13 known, active carbonic anhydrases (CAs), which differ in tissue distribution, subcellular
localization, kinetics, and sensitivity to various carbonic anhydrase inhibitors (Kaunisto et al., 2002; Leppilampi et
al., 2002; Parkkila et al., 2002; Sly and Shah, 2001; Winum et al., 2003). CA II is one of the most widespread of
Received 15 December 2003; accepted revised manuscript 26 May 2004.
© 2004 WILEY-LISS, INC.
DOI: 10.1002/humu.9266