War. Res. Vol. 23, No. 12, pp. 1599-1602, 1989 0043-1354/89 $3.00+0.00
Printed in Great Britain. All rights reserved Copyright © 1989 Pergamon Press plc
RESEARCH NOTE
THE ROLE OF FORMATE IN THE ANAEROBIC BAFFLED
REACTOR
A. GROBICK1 and D. C. STUCKEY
Department of Chemical Engineering, Imperial College, London SW7 2AZ, England
(First received October 1988; accepted in revised form May 1989)
Abstract--The anaerobic baffled reactor (ABR) contains a mixed anaerobic culture segregated into
compartments. During pseudo-steady state runs, formate was detected in the first two or three
compartments, thereafter dropping off sharply. Under conditions of shock loading, formate was detected
in the reactor effluent, up to peak concentrations of 2500 mg/l. There are indications that formate may
play an important role as an intermediate in the anaerobic digestion process, and that its production may
contribute significantly to reactor stability.
Key words--anaerobic digestion, formate, reactor design, reactor stability
INTRODUCTION
Formic acid has been known for many years to be
produced by mixed cultures during anaerobic fermen-
tation (Hungate et al., 1967; Marty, 1984). It is also
well known that there are many species of
methanogens which utilize formate directly to pro-
duce methane (Schauer et al., 1982; Daniels et al.,
1984). A new model of interspecies electron transfer
has now been proposed (Thiele and Zeikus, 1988)
which shows formate transfer to be far more signifi-
cant than hydrogen transfer, for selected species of
micro-organisms. In this paper we present experimen-
tal evidence of formate production and utilization, by
a mixed culture, in an anaerobic baffled reactor
(ABR). This is a novel type of reactor first described
by Bachmann et al. (1983).
EXPERIMENTAL DATA
The experiments reported in this paper were run on two
similar anaerobic baffled reactors of l0 I. reactor volume
(see Fig. 1), referred to as reactor 3 and reactor 4. Note that
the numerous gas offtakes are for research purposes only:
the basic design has a single undivided gas space. The
reactors were seeded with digester sludge from a sewage
treatment works. After the start-up period the micro-organ-
isms were observed to have flocculated. The feed composi-
tion (for l0 l.) was 26.67 g sucrose, 8 g peptone, 2.67 g Lab
Lemco, 0.8 g K2HPO 4 and 25 g NaHCO 3. The total COD
concentration was 4 g/1.
In addition to the standard analyses (COD, total sus-
pended solids) the bulk liquid was centrifuged, deep-frozen
and analysed by high pressure liquid chromatography
(HPLC). Liquid samples of 20 ml were withdrawn from
each compartment at steady state, via sampling tubes of
adjustable length set in the reactor cover. Sampling took
place from the last compartment back towards the first,
maintaining anaerobic conditions throughout. During
shock loading, samples were taken from the reactor effluent
only. The gas production rate was measured electronically,
but the gas composition was not monitored, due to the
difficulties of measuring and analysing the gas output from
16 compartments simultaneously.
Formate was observed in the anaerobic baffled reactors,
both at pseudo-steady state, and during shock loading. At
steady state (Figs 2 and 3), there was no formate detectable
in the effluent, but in the analyses of bulk liquid samples
from the individual compartments it appeared in the first
two or three compartments. The concentration of formate
was highest in the first compartment (1833 mg/l, reactor 3),
dropping off sharply thereafter.
Figures 4 and 5 show analyses of the effluent from the two
reactors at intervals of 0.5h, during shock load (101/h
flowrate) and on returning to the normal loading at 20 h
HRT (0.5 l/h). The shock loading took place from 0 to 3 h.
In both reactors the formate concentration rises rapidly,
peaks at 4 h (2466 mg/1, reactor 3) and disappears after 11 h.
The concentration of acetate increases rapidly, remains high
up to 7 h, then drops off. However, propionate builds up
rather slowly, remains high up to I 1 h, then drops rapidly.
Figure 6 shows the total COD and the COD contributed by
the VFAs for reactor 3. The total COD contributed by the
individual VFAs rises rapidly when the shock load begins,
remains high until 13 h, and then drops off to a very low
level.
DISCUSSION
Using the standard method of VFA analysis, an
FID gas chromatograph, formic acid is difficult to
detect as the peak is very close to the solvent peak.
Many workers prepare samples for analysis by acid-
ifying with formic acid (e.g. Cohen et al., 1980). Wet
chemical methods to determine formic acid in a
mixture of volatile fatty acids are lengthy (Jorgensen,
1981). This may explain why formate has not been
widely noted in the literature as a significant interme-
diate in the anaerobic digestion process.
The ABR used in this experimental work is a
unique design in that the gas space is partitioned.
This may give rise to an atmosphere in a given
compartment which is unusual for anaerobic di-
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