A Potential Synergistic Anticancer Effect of Paclitaxel and
Amifostine on Endometrial Cancer
Donghai Dai,
1
Anna M. Holmes,
1
Tan Nguyen,
1
Suzy Davies,
1
Daniel P. Theele,
2
Claire Verschraegen,
3
and Kimberly K. Leslie
1
1
Reproductive Molecular Biology Laboratory, Division of Maternal-Fetal Medicine, Department of Obstetrics and Gynecology;
2
Animal Research Facility, Department of Neuroscience; and
3
Cancer Research and Treatment Center, University of
New Mexico Health Sciences Center, Albuquerque, New Mexico
Abstract
Although paclitaxel is one of the most effective chemothera-
peutic agents, its usefulness is still limited in advanced and
recurrent endometrial cancer. Amifostine protection of
normal tissues against the side effects of chemotherapeutic
agents has been clinically proven in cancer patients; however,
its application in endometrial cancer has not been fully
evaluated. We have investigated the use of paclitaxel and
amifostine in controlling the growth of poorly differentiated
endometrial cancer cells, Hec50co, in vitro and in vivo . Our
studies show that amifostine had direct anticancer effects on
endometrial cancer cells in vitro by arresting the cell cycle
at the G
1
phase and inducing apoptosis. Amifostine also
inhibited s.c. tumor growth in athymic mice. Paclitaxel IC
50
value was reduced from 14 to 2 nmol/L with pretreatment of
a single dose of 178 Mmol/L of amifostine for 72 hours.
Amifostine also synergized with paclitaxel in the arrest of
the cell cycle at the G
2
-M phase and in the induction of
apoptosis. This two-drug regimen inhibited s.c. tumor growth
as well as improved mouse survival significantly more than
paclitaxel alone. Amifostine also significantly improved
paclitaxel-induced cytotoxic effects on peripheral blood
profiles. Our studies show that amifostine has direct
anticancer effects on endometrial cancer. Our data have
also shown a potential anticancer synergy between amifos-
tine and paclitaxel in vitro and in vivo, whereas amifostine
maintained a protective role in peripheral blood profiles.
The dual specificity of amifostine action should be further
investigated. (Cancer Res 2005; 65(20): 9517-24)
Introduction
Endometrial cancer is the fourth most common malignancy in
women (1). The American Cancer Society estimated that there
were 40,320 new cases and 7,090 deaths from endometrial cancer in
2004 (1). Currently, there is no effective therapy for advanced or
recurrent endometrial cancer.
Paclitaxel is an antimicrotubule agent which is among the most
potent single chemotherapeutic agents. Paclitaxel binds to the
h-tubulin subunit and stabilizes the microtubules, resulting in
disruption of normal microtubule dynamics during cell division (2).
Failure of microtubule separation during the G
2
-M phase blocks
cell mitosis and results in apoptosis. Application of this agent in
endometrial cancer treatment is under investigation. A Gyneco-
logic Oncology Group trial reports an overall response rate of 35.7%
in advanced or recurrent endometrial cancer (3). The response rate
is about 27.3% when paclitaxel is used in patients who have failed
prior chemotherapy (4). The overall survival, objective response,
and progression-free survival were improved by adding paclitaxel
to the doxorubicin and cisplatin regimen (5). However, the median
survival of patients treated with paclitaxel-based regimens is
around 12 months. Unfortunately, the benefits of paclitaxel are also
associated with serious adverse effects. These commonly include
myelosuppression, gastrointestinal toxicity, and peripheral neurop-
athy, among others (5). It was reported that 62% of patients treated
with paclitaxel had life-threatening leucopenia (3). In another
study, where paclitaxel was used as a second-line chemotherapy (4),
serious neutropenia occurred in more than 58% of patients with
one treatment-related death. Improvement of outcome with the
addition of paclitaxel to the doxorubicin and cisplatin regimen was
accompanied by a much higher treatment-related death rate (5).
Thus, it is imperative to search for more effective and safer
chemotherapeutic regimens.
Amifostine (WR2721) was developed in 1979 as an antiradiation
agent through the screening of 4400 chemicals by the Walter Reed
Army Institute of Research (6). Since then, its protective effects
against chemotherapy and radiotherapy in cancer patients have led
to amifostine being FDA-approved as a cytoprotective agent
against the adverse effects of chemotherapy and radiotherapy in
normal tissues. Amifostine is a prodrug, which must first be
converted to its active form, WR1065, by the enzyme alkaline
phosphatase, which is located on the cell membrane. WR1065 is
further metabolized through intracellular oxidation to the sym-
metrical disulfide WR33278, cysteamine, and mixed disulfides (7).
The underlying mechanisms of amifostine protective effects are
still not fully understood, but they are attributed to its ability to
scavenge free radicals (8) and to its antimutagenic effects (9). This
protection is based on the conversion of amifostine to the active
form, WR1065, which is rapidly taken up into normal tissues (10).
Amifostine dephosphorylation by membrane alkaline phosphatase
occurs effectively in normal tissue, which has a suitable pH of 6.6
to 8.2 for alkaline phosphatase to work. The uptake of WR1065 is
minimal in cancer cells because tumor tissue usually contains less
membrane-bound alkaline phosphatase (11–14) and the environ-
ment of the tumors is more acidic, making the enzymatic activity
less than optimal (15). Additionally, normal tissues contain the
WR1065 transporter that allows active absorption against a
concentration gradient (16). Taken together, these differences lead
to a concentration of WR1065 that is about 50 to 100 times higher
in normal tissue than in malignant tissue (16), thereby providing
normal tissues with a remarkable protection against cytotoxic
therapies.
Requests for reprints: Donghai Dai, Department of Obstetrics and Gynecology,
University of New Mexico Health Sciences Center, MSC 10 5580, Albuquerque, NM
87131. Phone: 505-272-0095; Fax: 505-272-3921; E-mail: DDai@salud.unm.edu.
I2005 American Association for Cancer Research.
doi:10.1158/0008-5472.CAN-05-1613
www.aacrjournals.org 9517 Cancer Res 2005; 65: (20). October 15, 2005
Research Article
Research.
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