Journal of Apicultural Research 45(3): 219–220 (2006) © IBRA 2006
NOTES AND COMMENTS
Evaluation of Beauveria bassiana
(Balsamo) Vuillemin (Deuteromycota:
Hyphomycetes) strains isolated from
varroa mites in southern France
W G Meikle
1*
, G Mercadier
1
, V Girod
2
, F Derouané
1
and W A Jones
1
1
European Biological Control Laboratory, USDA – ARS, Campus International de Baillarguet, CS 90013 Montferrier sur Lez, 34988 St.
Gely du Fesc, France
2
ADAPRO LR-CRALR, Maison des Agriculteurs, Mas de Saporta CS 30012, 34875 Lattes, France
Received 30 March 2006, accepted subject to revision 16 May 2006, accepted for publication 16 August 2006.
* Corresponding author. Email: wmeikle@ars-ebcl.org
Keywords: Apis mellifera, Varroa destructor, Beauveria bassiana, Metarhizium anisopliae, bioassay
Introduction
Varroa mites (Varroa destructor Anderson and Trueman) are an
increasingly important pest of honey bees (Shaw et al. 2002). Several
species of entomopathogenic fungi have been found to infect varroa
mites in the laboratory (Shaw et al. 2002) and have been tested in
the field (Kanga et al. 2003). The objectives of this study were to
isolate and characterize a strain of hyphomycete fungus that occurs
naturally in domesticated bee colonies and is virulent against varroa.
A total of 112 hives in 12 apiaries were visited in the
departments of Gard, Aude, Hérault, and Pyrénées Orientales
in France during 2005. Varroa mites were sampled by placing a metal
tray underneath the hive, opening the hive and pouring approximately
150 g of powdered sugar (Sucre Union, Paris, France) into the brood
box, making an effort to divide it evenly among the spaces between
frames. The hives were then closed and about 20 min. later all the
material that had collected on the tray was poured into a sealed
plastic box. The boxes were transported to the European Biological
Control Laboratory (EBCL), and all varroa mites were placed on
water agar (6 g/L) with chloramphenicol (0.4 g/L) to reduce bacterial
growth, incubated for 2 weeks at 23°C and observed for at least 3
weeks for sporulation. Entomopathogenic fungi collected in this way
were used to inoculate 2 types of media: Sabouraud dextrose agar
with yeast (SDAY) (Goettel and Inglis,1997) and semi-synthetic INRA
media (cf. Meikle et al. 2005), both with chloramphenicol (0.5g/L), in
plastic petri dishes (3.5 cm diam.). The most vigorous cultures were
re-isolated two to three times and then grown as pure cultures.
Two isolates from this exploration, as well as five isolates of
Metarhizium anisopliae (Metschnikoff) Sorokin (Deuteromycota:
Hyphomycetes) from the collection of entomopathogenic fungi at
EBCL, were evaluated in bioassays. Cultures of candidate isolates
were grown on SDAY with chloramphenicol and harvested after
12–14 days. Four experiments were conducted, each experiment
including two fungal isolates and the control treatment. Prior to
each experiment, conidial viability was assessed by plating a
suspension sample onto SDAY,incubating the plates at 23ºC for
24 h, and examining 200 conidia for germ tubes under a light
microscope. Conidial viability always exceeded 90%.
The day of the bioassay,white bee pupae were collected from hives
at EBCL by cutting sections of brood comb from the frame, placing the
pieces in a sterile plastic box and, in the lab, carefully extracting the
pupae and placing them individually in small (2x2x1 cm) plastic boxes
with screen covers. Any varroa mites found on the pupae upon their
removal from the comb were destroyed. Varroa mites used in the
bioassays were collected en masse the day of the bioassay using the
powdered sugar treatment described above.
While the mites and bee pupae were being collected, a 10
7
conidia/ ml suspension was prepared by adding conidia to 10 ml
of a 0.1% aqueous solution of Tween 80®, pouring the mixture
into a 50 ml bottle filled with glass beads, and agitating the bottle
for 2 min. using a Vortex Genie 2 (Scientific Industries, Bohemia,
NY, USA). Thirty varroa mites were then placed in a petri dish
(9 cm. diam.) lined with moist filter paper. Care was taken to
place mites as far as possible from each other. The dish was
placed in a spray tower (Burkard Manufacturing Ltd., Herts.,
England) and sprayed with either an aqueous Tween 80®
solution (control), or the conidial suspension. The spray tower
had an air pressure of 0.74 kg/cm
2
and delivered on average 714
conidia/ mm
2
using suspensions of 10
7
conidia/ ml (Meikle et al.
2005). After exposure, two varroa mites were placed on each