JOURNAL OF THE
WORLD AQUACULTURE SOCIETY
Vol. 41, No. 3
June, 2010
Effect of Dietary Carotenoids on Skin Color and Pigments
of False Clownfish, Amphiprion ocellaris, Cuvier
Inayah Yasir
Department of Marine Sciences, Hasanuddin University, Makassar 90245 Indonesia
Jian G. Qin
1
School of Biological Sciences, Flinders University, GPO Box 2100,
5001 SA Adelaide, Australia
Abstract
This study evaluated the role of supplemented dietary carotenoids in regulating the skin color and
pigments of the false clownfish, Amphiprion ocellaris. Three carotenoid types, such as astaxanthin,
β-carotene, and canthaxanthin, were added to the basal diet at four pigment doses (0, 20, 50,
and 100 ppm). Carotenoid diets were feed for 5 wk and then withdrawn from the diet for three
additional wk during an 8-wk trial. The dose of each diet did not change the overall color hue,
brightness, or saturation, but astaxanthin was the only carotenoid that enhanced red hue by the
end of Week 5. The withdrawal of astaxanthin from the diet did not reduce the red hue, but
reduced saturation. In contrast, the withdrawal of dietary β-carotene or canthaxanthin reduced
color saturation and brightness, but did not affect color hue. Dietary astaxanthin increased skin
astaxanthin in Week 1 and skin zeaxanthin in Week 5. The withdrawal of astaxanthin escalated
skin canthaxanthin and zeaxanthin by Week 8. Dietary β-carotene suppressed skin β-carotene,
but enhanced skin zeaxanthin by Week 8. Although skin canthaxanthin was enhanced by dietary
β-carotene from Week 5 onward, dietary β-carotene at 100 ppm maximized skin canthaxanthin by
Week 8. Interestingly, dietary canthaxanthin suppressed skin canthaxanthin and zeaxanthin, but
increased β-carotene. This study suggests that astaxanthin has the potential to enhance the red hue
on clownfish skin and its withdrawal from the diet did not fade the red hue of the skin.
The false clownfish, Amphiprion ocellaris,
has been commercially produced for aquarium
hobbyists (Hoff 1996; Chapman et al. 1997).
A noble reason for culturing clownfish is to
protect coral reefs from destructive collection
and over exploitation of the wild clownfish
population (Alava and Gomes 1989; Andrews
1990). Although the culture of clownfish has
been successfully achieved (Alava and Gomes
1989; Maroz and Fishelson 1997), the col-
oration of farm-raised fish is inferior to their
wild counterparts (Booth et al. 2004). The color
of wild-caught clownfish is usually bright pink
or reddish orange, while hatchery-raised fish
are light orange (Tanaka et al. 1992). A recent
study reported that a light intensity of 20–50 lx
1
Corresponding author.
could brighten the skin color of clownfish (Yasir
and Qin 2009a) and this color could add to
the commercial value in the aquarium trade
(Hoff 1996). Yasir and Qin (2009b) further
demonstrated that a blue or green background
could strengthen the orange color, whereas a
white background made fish less color saturated
but brighter. Our previous studies showed that
clownfish could temporarily change coloration
through changing light intensity or background
color. However, manipulating environmental
conditions is unlikely to permanently alter the
pigment composition of fish skin. An alterna-
tive is to explore the response of coloration of
clownfish to diet manipulation.
Carotenoids are biosynthesized by plants,
algae, and certain yeast and bacteria (Ong
and Tee 1992). Carotenoids are responsible for
the red, orange, and yellow colors of fish and
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