ABSTRACT
Sea floor mapping along the Apulia continental shelf (Italy)
verified the abundance of autochthonou red algae build-ups, mapped
as coralligenous habitats (CHs), in a water depth range of 5-100 m. In
general, CHs were found to develop three dimensional structures,
with a rigid framework and to represent an important geomorpho-
logical and sedimentological element on the Mediterranean shelf.
Here, we provide the first geomorphological description of CHs
(thus poorly categorized) using acoustic data obtained from Side
Scan Sonar (SSS) and MultiBeam (MB) echosounder surveys,
ground-truthed using a ROV and underwater camera. In SSS
mosaics, CHs generally yielded intermediate to high backscatter in
response to a rigid cavernous framework. According to the various
shapes and the lateral continuity that coralligenous build-ups dis-
played in explored locations, two distinct textures were determined to
be present. Various geomorphological expressions of CHs were noted
within our dataset and in images obtained from MB bathymetry. We
determined that coralligenous build-ups are typically represented by
positive-relief structures that vary from isolated blocks (randomly
scattered on a generally flat mobile soft bottom) to a field of blocks
(adjacent or even coalescent), and/to ridge with several metres of
lateral continuity. In most cases, CHs occurred on flat mobile soft
bottom, thus representing an example of coralligenous de plateau.
Our results characterize for the first time the CHs through
seafloor mapping techniques, which demonstrated to represent an
instrumental tool for their geomorphological characterization.
KEY WORDS: Coralligenous, geomorphology, habitat map-
ping, Mediterranean Sea.
INTRODUCTION
Modern autochthonous red algae build-ups, corallige-
nous build-ups, are a well-known feature of the temperate
Mediterranean Sea (FELDMANN, 1937; PÉRÈS & PICARD,
1964; LABOREL, 1961; LAUBIER, 1966; PÉRÈS, 1967;
HONG, 1982; BOSENCE, 1983, 1985; SARTORETTO, 1994;
SARTORETTO et alii, 1996; DI GERONIMO et alii, 2002;
BALLESTEROS, 2006; BEN MUSTAPHA et alii, 2003; CASEL-
LATO & STEFANON, 2008; AGNESI et alii, 2009). Due to
their high level of biological diversity (PÉRÈS &PICARD,
1964; BOSENCE, 1983; BALATA et alii, 2005; BALLESTEROS,
2006), they are listed in the EC Habitat Directive.
In the framework of Mediterranean marine benthic
zonation, coralligenous habitat (CHs) are indicative of a
circalittoral biocoenosis consisting of a three dimensional
biogenic build-up that forms a solid substrate primarily
dominated by coralline algae (PÉRÈS & PICARD, 1964;
BELLAN-SANTINI et alii, 1994; BRESSAN et alii, 2001). Due
to their three-dimensional structure, CHs often play a
primary role in the geomorphological development of
the continental shelf.
CHs develop on both hard and soft bottoms, in 4 to
160 m of water depth (wd) (SARÀ, 1966; FREDJ, 1964;
LABOREL, 1961, 1987; SARTORETTO, 1994; BALLESTEROS,
2006) and adapt to low levels of nutrients and tempera-
ture from 10 to 23°C in waters with moderate hydro-
dynamics (BOSENCE, 1983; BALLESTEROS, 2006). The
extreme variability of their bathymetrical distribution is
partially governed by the light-dependency of participat-
ing benthic organisms (BALLESTEROS, 1992; MARTÌ et alii,
2004), a finding particularly true for macroalgae, adapted
to grow at dim-light conditions (LITTLER et alii, 1986;
VADAS & STENECK, 1988).
Architecture and morphology are primarily controlled
by biological carbonate productivity that responds to cli-
mate, oceanography, physiography, changes in accommo-
dation space and terrigenous supply (SCHLAGER, 1991,
1993; BETZLER et alii, 1997).
Modern Mediterranean coralligenous build-ups are
characterized by large structures that may be up to 4 m
high and greater than 50 m in lateral continuity
(BOSENCE, 1983, 1985).
Coralligenous build-ups vary in shape and dimension:
nevertheless, their geomorphological expression have not
been exhaustively categorized. Various definitions for
characterizing CHs structures, reflecting different con-
structional morphologies, have been reported in the scien-
tific literature. For examples, CHs have been identified
as coralligenous de plateau (BOSENCE, 1985), columnar
crustose coralline algal build-ups (DI GERONIMO et alii,
2001a,b, 2002), and algal reefs (BOSENCE, 1983) or banks
(BALLESTEROS, 2006). Definitions are generally based on
scuba diver’s visual observations or direct sampling. From
a geomorphological point of view, the following main
morphologies have been distinguished: 1) banks-flat
frameworks with thickness ranging from 0.5 to 4 m
mainly built over more or less horizontal substrata, and
2) rims-structures on submarine vertical cliffs or sur-
rounding the opening of submarine caves, generally
located in shallower waters than banks (PÉRÈS & PICARD,
1964; LABOREL, 1987; BALLESTEROS, 2006). If rims
develop on a hard original substrate, banks are generally
reported as a consequence of the coalescence of rhodoliths
(PÉRÈS & PICARD, 1952), indicated as a coralligenous de
plateau (PÉRÈS & PICARD, 1964). However these frame-
(*) Università di Milano-Bicocca, Dipartimento di Scienze
dell’Ambiente, del Territorio e di Scienze della Terra. Piazza della
Scienza, 4 - 20126 Milano, Italy.
Coralligenous habitat in the Mediterranean Sea:
a geomorphological description from remote data
VALENTINA BRACCHI (*), ALESSANDRA SAVINI (*), FABIO MARCHESE (*), SERENA PALAMARA (*),
DANIELA BASSO (*) & CESARE CORSELLI (*)
251-R1 – BRACCHI ET ALII
Ital. J. Geosci. (Boll. Soc. Geol. It.), Vol. 134, No. 1 (2015), pp. 00-00, 8 figs. (doi: 10.3301/IJG.2014.16)
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