African Journal of Biotechnology Vol. 10(18), pp. 3521-3527, 2 May, 2011
Available online at http://www.academicjournals.org/AJB
DOI: 10.5897/AJB10.1389
ISSN 1684–5315 © 2011 Academic Journals
Full Length Research Paper
Maize breeding for marginal lands: Physiological and
molecular approach to decipher response and selection
of maize recombinant inbred lines (RILs) under water
deficit at early growth stage
Malik Nawaz Shuja
1
, Wajid Ali
1
, Aqib Iqbal
1
*, Ijaz Ali
1
, Iqbal Munir
1
, Dawood Ahmad
1
,
Inamullah
2
, Shaheenshah
2
, Gulzar Ahmad
3
, M. Arif Khan
2
and Zahoor Ahmad Swati
1
1
Laboratory of Molecular Genetics, Institute of Biotechnology and Genetic Engineering, Agricultural University,
Peshawar, Khyber Pakhtoonkhwa, Pakistan.
2
Department of Agronomy, Agricultural University, Peshawar, Khyber Pakhtoonkhwa, Pakistan.
3
Cereals Crops Research Institute, Pirsabak, Nowshera, Pakistan.
Accepted 3 January, 2011
Drought is the major abiotic stress constraining the production of maize (Zea mays L.) in the arid and
semi-arid areas. Recombinant inbred lines (RILs) with improved performance from the 6th selection
cycles were evaluated against drought stress at early growth stages and genetic distance was
determined through random amplified polymorphic DNA (RAPD) primers. Significant variation was
observed in the rate of water loss (RWL) at 30, 90 and 150 min after excision, relative water content
(RWC) and membrane stability among the RILs. In comparison with check variety Azam, 7 RILs
maintained a low RWL, 2 had higher RWC and 6 RILs had higher membrane stability, thus showing a
higher degree of adaptation to drought stress. The preliminary results showed polymorphism among
the drought sensitive and tolerant RILs. The RILs were clustered into three groups on the basis of
amplification pattern obtained with 60 RAPD markers. Variation in the genetic makeup of the tolerant
RILs was evident from their clustering in different groups, though most were clustered in group III.
Furthermore, one RAPD marker could identify maize genotypes, maintaining low RWL from the excised
leaves. We are in the process of sequencing the amplified product of this primer and transforming it
into sequence characterized amplified region (SCAR) markers for a more reliable marker assisted
selection.
Key words: Maize recombinant inbred lines (RILs), drought, physiological markers, random amplified
polymorphic DNA (RAPD).
INTRODUCTION
Global climate change is now generally considered to be
underway and is expected to result in a long-term trend
towards higher temperatures, greater evapotranspiration
and an increased incidence of drought in specific regions
( Hillel and Rosenzweig, 2002). Under these conditions,
research into plant breeding for developing improved
crop varieties and management practices to enhance
*Corresponding author. E-mail: aqib72@aup.edu.pk. Tel: +92-
336-9086080. Fax: +92-91-9218102.
plant water use efficiency and growth, when water is limi-
ting has become increasingly important. As one of the
major abiotic stress, drought causes differences between
the mean and potential yield, as well as variation from
year to year. It has been estimated that crops attain only
25% of their potential yield because of the detrimental
effects of environmental stresses (Boyer, 1982). Maize is
cultivated worldwide under very diverse climatic condi-
tions and has the largest total annual grain production in
the world (590.5 million metric tons, mmt) among the
major grain crops. Though maize is essential for global
food security, in several key production environments, the