~ Pergamon
0306-4522(95)00524-2
Neuroscience Vol. 72, No. 2, pp. 505-518, 1996
Elsevier Science Ltd
Copyright © 1996 IBRO
Printed in Great Britain. All rights reserved
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INCREASED EXPRESSION OF TYPE 1 INSULIN-LIKE
GROWTH FACTOR RECEPTOR MESSENGER RNA IN
RAT HIPPOCAMPAL FORMATION IS ASSOCIATED WITH
AGING AND BEHAVIORAL IMPAIRMENT
K. L. STENVERS,* P. K. LUND*t~ and M. GALLAGHER*§
*Curriculum in Neurobiology and Departments of ~'Physiology, :~Pediatrics, and §Experimental
Psychology, University of North Carolina, Chapel Hill, NC 27599, U.S.A.
Abstract--Insulin-like growth factor messenger RNAs are expressed in adult rat brain. However, little is
known about the effects of aging on the expression of the insulin-like growth factors, their receptors, and
their binding proteins in different regions of rat brain. The goal of the current study was to assess whether
there is altered expression of the insulin-like growth factor system during normal aging in the hippocampal
formation, a region particularly vulnerable to the aging process. A spatial learning task in the Morris water
maze was used to assess the cognitive status of young (7-8-month-old) and aged (28 29-month-old) male
Long-Evans rats. Sites of expression and abundance of insulin-like growth factor-I, type 1 insulin-like
growth factor receptor, and insulin-like growth factor binding protein-4 messenger RNAs were then
examined by in situ hybridization histochemistry and solution or northern blot hybridization assays. In
situ hybridization histochemistry revealed no qualitative differences in the regional distribution of
insulin-like growth factor-I, type 1 receptor, and insulin-like growth factor binding protein-4 messenger
RNAs within the hippocampal formation of young and aged rats. However, quantitative analysis of
messenger RNA abundance in hippocampal tissue homogenates showed a significant age-related increase
in type I receptor messenger RNA (n = 25; t = -2.5; P < 0.02). Furthermore, linear regression analysis
indicated that type 1 receptor messenger RNA abundance was significantly correlated with spatial learning
impairment in the water maze (r = 0.44; P < 0.03) such that greater behavioral impairment was associated
with higher type 1 receptor messenger RNA levels in the hippocampal formation. Neither insulin-like
growth factor-I nor insulin-like growth factor binding protein-4 messenger RNA abundance was related
to age or behavior. However, linear regression revealed a negative correlation between insulin-like growth
factor-I messenger RNA abundance and type 1 receptor messenger RNA abundance in aged hippocampus
(r = -0.72, P < 0.01).
These data indicate that increased hippocampal expression of type 1 receptor messenger RNA is
associated with aging and cognitive decline. The correlation between type 1 receptor and insulin-like
growth factor-I messenger RNA abundance in the hippocampal formation of aged rats suggests that
insulin-like growth factor availability may influence type 1 receptor expression. However, because no
overall age difference was found in the amount of insulin-like growth factor-I messenger RNA in the
hippocampal formation, decreased insulin-like growth factor from other sources such as the cerebrospinal
fluid and the peripheral circulation may be involved in up-regulating type 1 receptor messenger RNA.
Alternatively, type 1 receptor messenger RNA regulation may be part of atrophic response to the
degenerative and regenerative events that occur within the hippocampal formation during aging.
Key words: insulin-like growth factor-I, insulin-like growth factor binding protein-4, in situ hybridization
histochemistry, northern blot, solution hybridization, spatial learning.
A considerable body of evidence indicates that hippo-
campal circuitry is especially vulnerable to the neu-
rodegeneration associated with normal and
pathological aging. However, aged hippocampus ap-
pears to retain some of its capacity to regenerate in
response to this damage) 6'4°'65 A variety of morpho-
Abbreviations: ANOVA, analysis of variance; CA1 3,
subfields of Ammon's horn; cDNA, complementary
deoxyribonucleic acid; cRNA, complementary ribonucleic
acid; CTP, cytidine thiotriphosphate; EDTA, ethylenedi-
aminetetraacetic acid; IGF-I, insulin-like growth factor-I;
IGFBP-4, insulin-like growth factor binding protein-4;
NTF, neurotrop~hic factor; SSC, standard saline citrate;
UTP, uridine 5'-triphosphate.
logical changes, including axonal sprouting and
synaptic reorganization, has been reported in hippo-
campal regions sustaining loss of cells or input from
l e s i o n s , 24"33"74"76 disease, 1'33"46 and normal aging, z627
Neurotrophic factors (NTFs) are believed to be
necessary for such structural reorganization. 4°'65 Var-
ious NTFs and their receptors are synthesized locally
in the hippocampus. Little is known about the actions
of these NTFs in adult brain, although available data
suggest that they play important roles in maintaining
t h e v i a b i l i t y o f h i p p o c a m p a l n e u r o n s . 4'13'32'49'6°~61
Recent data suggest that one family of NTFs, the
insulin-like growth factors (IGFs), may have actions
on a broad spectrum of hippocampal cells. 4"7A3'17"49"75
NSC 72/2--H
505