        !" #$ % &                    !"#$ " %  &&’ (")$ " *+ ,*-.* ( " /0 0* (1 1 Medical Research Council Functional Genomics Unit, Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford OX1 3PT, United Kingdom 2 Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford OX1 3QX, United Kingdom 3 Centre for Biomedical Research, Facultad de Medicina, Universidad Diego Portales, Santiago 8370068, Chile 4 Centre for Clinical Brain Sciences, The University of Edinburgh, Scotland, United Kingdom *Correspondence: zoltan.molnar@dpag.ox.ac.uk $ Both authors contributed equally to this work 2 3*+& Intermediate progenitors, progenitors, ventricular radial glia, subventricular zone, subventricular (outer) radial glia, transcriptomics, cerebral cortex, Wulst, Dorsal ventricular ridge. Review The Journal of Comparative Neurology Research in Systems Neuroscience DOI 10.1002/cne.23871 This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process which may lead to differences between this version and the Version of Record. Please cite this article as an ‘Accepted Article’, doi: 10.1002/cne.23871 © 2015 Wiley Periodicals, Inc. Received: Apr 03, 2015; Revised: Jul 21, 2015; Accepted: Jul 21, 2015 This article is protected by copyright. All rights reserved.