Pure & Appl.Che,j., Vol.54, No.10, pp.l935—I950, 1982. 00334545/821b0193516$03.00/0
Printed in Great Britain.
Pergamon Press Ltd.
©1982 IUPAC
STEROIDAL GLYCOSIDES FROM STARFISHES
L. Minale , C. Pizza, R. Riccio and F. Zollo
Istituto di Chimica Biorganica, Università, Via RodinO, 22,Napies,
and Istituto di Chimica di Molecole di Interesse Biologico del
C.N.R., Arco Felice, Naples, Italy
Abstract -
This paper covers recent work - much of it from the
author's laboratory -
dealing
with novel steroidal oligoglyco-
sides from starfishes. The steroidal glycosides until now
encountered in this class of marine animals can be grouped
into three structure types. Compounds of the first type, reco-
gnized for long time, include the sulphated saponins (astero-
saponins) ,
characterized
by steroidal aglycones possessing a
3,6ct-diol pattern and a 9,11-double bond; the oligosaccharide
moiety (four up to six sugar units) is attached at C-6 and the
sulphate residue is at C-3. Compounds of the second type,
recently discovered in two species of the genus Echinaster,
have a number of unusual features: a L7,3,613-dihydroxy stero-
idal moiety, there is no sulphate group and, most remarkably,
the carbohydrate chain (three sugar units) is cyclized between
C-3 and C-6 of the aglycone. Compounds of the third type
include glycosides having highly hydroxylated steroidal agly-
cones; the carbohydrate moiety (one or two sugar units) is
attached at C-24 of the side chain and there is no sulphate
residue. During the course of our investigation on the stero-
idal glycosides from starfishes we have also encountered
several polyhydroxylated sterols and their structures are
presented.
INTRODUCTION
Saponins, water soluble compounds composed of sugars and steroid or triterpen-
oid moieties, are common constituents of terrestrial plants, but are uncommon
as animal constituents. In the animal kingdom saponins have been found
in the exclusively marine phylum Echinodermata and particularly in species of
the classes Holothuroidea (sea cucumbers) and Asteroidea (starfishes) . These
compounds are apparently absent from the other three classes, Crinoidea (sea
lilies), Echinoidea (sea urchins) and Ophiuroidea (brittle stars), of echino-
derms.
Saponins derived from sea-cucumbers (holothurins) are triterpenoid glycosides,
which, upon acid hydrolysis, give triterpenoid aglycones based on the lanos-
tane skeleton, sugars and sulphate, whereas those from starfishes (asterosa—
ponins) are sulphated steroidal glycosides (Ref s. 1, 2, 3).
The toxicity of starfishes has been described in various ways for many years,
but most of the reports can be explained by the presence of saponins. Starfish
extracts and also purified saponins have been reported to exhibit a broad
spectrum of physiological and pharmacological activities. Asterosaponins are
highly surface active, and most of them show potent haemolitic properties
(Ref s. 2, 4). They also have antitumor (Ref. 5) and antibacterial activities
(Ref. 6); Shimizu reported that asterosaponins from Asterias forbesi,
ster planci and Asterina pectinifera inhibited influenza virus multiplication
(Ref. 7). Antiinflammatory activity was reported for Asterias forbesi saponins
(Ref. 8), and activity toward contraction of the rat phrenic nerve-diaphram
preparation was reported for Asterias amurensis saponins (Ref. 9). Several
observations concerning with the biological functions of the asterosaponins
have also been described. Because of their general toxicity it is probable
that saponins act primarily as chemical defence agents, discouraging many
predators; they also induce escape reactions in bivalve molluscs (Ref. 10),
thus reducing the predation ability of the starfishes theirself.
Asterosaponins have been identified as the spawning inhibitor in the Japanese
starfish Asteria amurensis (Ref. 11) and more recently they have been reported
1935