Journal of Coastal Research SI 63 x-xx Coconut Creek, Florida 2013
DOI: 10.2112/SI63-001.1 received (Day) (Month) 2012; accepted
(Day) (Month) 2012.
© Coastal Education & Research Foundation 2013
www.JCRonline.org
www.cerf-jcr.org
Seasonal summer stratiication and enhanced nutrient loading of the Louisiana continental shelf (USA) west of the Mississippi
River create hypoxic regions that affect large areas of the benthos. Total sediment oxygen uptake and nutrient recycling were
measured during different, (e.g. pre-, early, late and post-) hypoxic regimes using shipboard Batch Micro-Incubation Chambers
(BMICs) in 2004 to 2005 and again in 2007 to 2009. Sediment community oxygen consumption during oxic regimes (dissolved
oxygen > 63 µmol L
-1
) was -9.5 ± 0.7 mmol O
2
m
-2
d
-1
(mean ± SE), almost twice that measured (-5.8 ± 0.6 mmol O
2
m
-2
d
-1
)
during suboxic conditions. During the summer when hypoxia occurred, the benthos consumed nitrate and nitrite (-0.14 ± 0.04 and
-0.10 ± 0.02 mmol N m
-2
d
-1
respectively) and produced ammonium (1.6 ± 0.39 mmol N m
-2
d
-1
). Elevated sediment community
oxygen consumption and nutrient remineralization occurred near terrestrial river inputs associated with the Mississippi and
Atchafalaya Rivers. Net release of dissolved inorganic nitrogen, in the form of ammonium, peaked during late summer. Released
ammonium may be a source of nutrients for primary production in bottom waters, and can also provide reduced nitrogen for
nitriication and microbial respiration, both of which may reinforce the intensity and duration of hypoxia. Based on chamber
results, sediments actively scavenged phosphate from the bottom waters (-98.4 ± 21.3 µmol P m
-2
d
-1
) and released silicate (2.62
± 0.31 mmol Si m
-2
d
-1
). The addition of reactive nitrogen and removal of phosphorous due to benthic community metabolism
could potentially be accentuating phosphorous limitation on the continental shelf.
ADDITIONAL INDEX WORDS: Louisiana continental shelf, Mississippi-Atchafalaya river system, sediment
biogeochemistry, remineralization, hypoxia, sediment oxygen demand, positive feedback.
†Department of Oceanography
Texas A&M University
3146 TAMU
College Station, Texas 77843-3146
‡Department of Marine Biology
Texas A&M University at Galveston
200 Seawolf Parkway
Galveston, Texas 77553
§University of Hawaii at Manoa,
Department of Oceanography
1000 Pope Road
Honolulu, HI 96822
cnunn@hawaii.edu
ABSTRACT
Nunnally, C.C.; Rowe, G.T..; Thornton, D.C.O., and Quigg, A.S., 2012. Sedimentary oxygen consumption and nutrient
regeneration in the Gulf of Mexico hypoxic zone. In: Brock, J.C., Barras, J.A., and Williams, S.J. (eds.), Understanding and
Predicting Change in the Coastal Ecosystem of the Northern Gulf of Mexico, Journal of Coastal Research, Special Issue, No.
63, pp. x-xx. Coconut Creek (Florida), ISSN 0749-0208.
Sedimentary Oxygen Consumption and Nutrient Regeneration in the
Northern Gulf of Mexico Hypoxic Zone
Clifton C. Nunnally†§, Gilbert T. Rowe†‡, Daniel C. O. Thornton†, and Antonietta Quigg†‡
INTRODUCTION
Under the Mississippi-Atchafalaya River plume, along the
Louisiana continental shelf (Figure 1), expansive low oxygen
areas form during summer months due to strong thermohaline
stratiication (Bianchi et al., 2010; Hetland and DiMarco, 2008;
Rabalais, Turner, and Wiseman, 1999; Turner et al., 2005;
Wiseman et al., 1997) and elevated riverine nutrient inputs
(Dortch et al., 1994; Lehrter, Murrell and Kurtz, 2009; Quigg
et al., 2011) and sediments (Rowe 2001, Rowe et al., 2002)
beneath the pycnocline. Over time and under strong seasonal
stratiication, this leads to large zones of hypoxia in bottom
waters that persist until oxygen can be physically mixed from
the surface or offshore. These processes of eutrophication,
stratiication, and respiration combine to create the northern
Gulf of Mexico Hypoxic Zone (GOMHZ). All of these processes
vary in time and space and thus variations in sedimentary
biogeochemical processes (Morse and Rowe, 1999; Rowe et al.,
2002) need to be understood to better predict consequences of
river runoff (Dagg and Breed, 2003; Quigg et al., 2011; Rabalais,
Turner, and Wiseman, 1999), wetland particulate and dissolved
organic carbon (POC and DOC) contributions (Bianchi et al.,
2009, 2010, 2011; Rosenbauer et al., 2009; Swarzenski et al.,
2008), and physical forcing (Cochrane and Kelly, 1986; DiMarco
et al., 2009; Wiseman et al., 1997).
River plume dynamics and coastal boundary currents create
unique zones with differing hypoxic potential within the GOMHZ
(Rowe and Chapman, 2002). Near the river mouth, a ‘brown’
zone of heavy sediment loading impedes light penetration and
limits primary productivity within the plume. Further away from
the river plume, a ‘green’ zone is deined by elevated nutrients
and high primary productivity, which ostensibly deposits large
loads of marine-derived organics to the benthos (Turner and
Rabalais, 1994). West of Terrebonne Bay (Figure 1), a ‘blue’
zone is dominated by intense seasonal stratiication and a strong
pycnocline; nutrients are limiting at this distance from river input
and most primary production is fueled by recycled nutrients
(Dortch and Whitledge, 1992; Quigg et al., 2011). Transition
areas associated with coastal bays (Timbalier, Terrebonne
and Atchafalaya Bays) between these zones provide inputs of
dissolved organic matter independent of the main Mississippi
River outlow (Bianchi et al., 2010).