Planning and Control of Sequential Rapid
Aiming in Adults With Parkinson’s Disease
A. L. Smiley-Oyen
K. A. Lowry
Motor Control and Learning Research Laboratory
Department of Health and Human Performance
Iowa State University, Ames
ABSTRACT. Eight people with Parkinson’s disease (PD), 8 age-
matched older adults, and 8 young adults executed 3-dimensional
rapid aiming movements to 1, 3, 5, and 7 targets. Reaction time,
flight time, and time after peak velocity to the 1st target indicated
that both neurologically healthy groups implemented a plan on the
basis of anticipation of upcoming targets, whereas the PD group
did not. One suggested reason for the PD group’s deficiency in
anticipatory control is the greater variability in their initial force
impulse. Although the PD group scaled peak velocity and time to
peak velocity similarly to the other groups, their coefficients of
variation were greater, making consistent prediction of the move-
ment outcome difficult and thus making it less advantageous to
plan too far in advance. A 2nd finding was that the PD group
exhibited increased slowing in time after peak velocity in the final
segments of the longest sequence, whereas the other 2 groups did
not. The increased slowing could be the result of a different move-
ment strategy, increased difficulty modulating the agonist and
antagonist muscle groups later in the sequence, or both. The
authors conclude that people with PD use more segmented plan-
ning and control strategies than do neurologically healthy older
and young adults when executing movement sequences and that
the locus of increased bradykinesia in longer sequences is in the
deceleration phase of movement.
Key words: anticipation, basal ganglia, motor planning, sequence,
variability
t is well established that people with Parkinson’s disease
(PD) exhibit difficulties with movement sequencing in such
tasks as handwriting (Phillips, Chiu, Bradshaw, & Iansek,
1995; Teulings, Contreras-Vidal, Stelmach, & Adler, 1997),
buttoning (Rocca, Maraganore, McDonnell, & Schaid, 1998;
Soliveri, Brown, Jahanshahi, & Marsden, 1992), keyboarding
(Freund, 1989; Longstreth, Nelson, Linde, & Munoz, 1992),
and even finger tapping (Camicioli, Grossmann, Spencer,
Hudnell, & Anger, 2001; Harrington, Haaland, & Her-
manowicz, 1998). People with PD move more slowly when
performing a movement sequence than when performing dis-
crete movements, and they exhibit longer pause times
between elements of the sequence (Agostino, Berardelli,
Formica, Accornero, & Manfredi, 1992; Benecke, Rothwell,
Dick, Day, & Marsden, 1987; Harrington & Haaland, 1991;
Weiss, Stelmach, & Hefter, 1997). They also exhibit an
increase in bradykinesia, that is, movement slowness, in the
later segments of a sequence (Agostino et al., 1992; Agostino
et al., 1994; Harrington & Haaland).
One reason that execution of movement sequences may
be different in PD is that they are planned differently. One
can measure planning on the basis of changes in reaction
time (RT, i.e., the time required to preprogram the move-
ment); as the number of elements in a rapidly executed series
of discrete movements increases, so does RT. Investigators
have concluded that as this type of sequence increases in
number of elements, the movement is more complex; and
therefore more neurological and psychological processing
time is required before the movement is initiated (Christina,
1992; Fischman, 1984; Garcia-Colera & Semjen, 1987; Gor-
don & Meyer, 1987; Henry & Rogers, 1960; Lajoie &
Franks, 1997; Sidaway, 1991; Sidaway, Christina, & Shea,
1988; Smiley-Oyen & Worringham, 2001; Sternberg, Mon-
sell, Knoll, & Wright, 1978). A longer RT for longer
sequences indicates that young adults plan a movement
sequence in an integrative fashion, taking into account many
segments, not just the first one or two segments.
People with PD exhibit increased preprogramming time
comparable with that of age-matched controls for two- and
Correspondence address: A. L. Smiley-Oyen, 244 Forker Build-
ing, Department of Health and Human Performance, Iowa State
University, Ames, IA 50011, USA. E-mail address: asmiley
@iastate.edu
103
I
Journal of Motor Behavior, 2007, Vol. 39, No. 2, 103–114
Copyright © 2007 Heldref Publications
J. P. Kerr
Voxel Research
Ames, Iowa