AQUATIC BIOLOGY
Aquat Biol
Vol. 7: 107–112, 2009
doi: 10.3354/ab00194
Printed November 2009
Published online October 22, 2009
INTRODUCTION
Horseshoe crab is the common name of 4 species of
marine arthropods (Xiphosura, Chelicerata) found on
the east coasts of Asia and North America. Despite
their name, these animals are not crabs at all, but are
related to scorpions, spiders and extinct trilobites.
Tachypleus tridentatus (Leach 1819), known as the
Japanese or Chinese horseshoe crab, is a species in the
family Tachypleinae.
Tachypleus tridentatus larvae tend to stay in their
nests immediately after hatching, where they usually
reside over winter. They then leave the nests and live
at or near their natal beaches throughout the next
spring or summer (Sekiguchi 1988). Juvenile T. triden-
tatus bury themselves in the sand/mud sediment dur-
ing high tide and emerge on the sediment surface to
feed during low tide (Chen et al. 2004, Chiu & Morton
2004); therefore, they can be found at the sediment
surface or buried in sediments in their nursery inter-
tidal zone (Rudloe 1979). Adult T. tridentatus spend the
winter in hibernation on the offshore ocean bottom at
depths of 20 m, and then migrate from their winter
deeper-water habitat to the shallow areas of coves
when seawater temperatures rise in the summer (Seki-
guchi 1988). The life-history characteristics and habitat
preferences of the Chinese horseshoe crab suggest
that the dispersal capability of juveniles might be
restricted, resulting in substantial population subdivi-
sions on a small geographic scale (Pierce et al. 2000,
King et al. 2005).
Populations of Tachypleus tridentatus in Asia have
declined sharply for the past several decades. In Japan,
T. tridentatus has been designated as a protected spe-
cies since 1928 owing to near extinction, but its popula-
tion size remains under threat (Botton et al. 1996). Hong
Kong populations have shown similar trends, with de-
clines in juvenile densities found on nursery beaches in
© Inter-Research 2009 · www.int-res.com *Corresponding author. Email: bhpshin@cityu.edu.hk
Summer distribution and abundance of juvenile
Chinese horseshoe crabs Tachypleus tridentatus
along an intertidal zone in southern China
Menghong Hu
1
, Youji Wang
1
, Yan Chen
1
, Siu-Gin Cheung
1
, Paul K. S. Shin
1,
*,
Qiongzhen Li
2
1
Department of Biology and Chemistry, City University of Hong Kong, Tat Chee Avenue, Kowloon, Hong Kong, SAR
2
Guangxi Institute of Fisheries, Nanning, Guangxi, 530021, PR China
ABSTRACT: Owing to human exploitation, pollution and loss of breeding and nursery grounds,
Chinese horseshoe crabs Tachypleus tridentatus are facing the risk of population decline and even-
tual extinction. In order to provide baseline information needed to formulate future conservation
strategies, we examined the spatial distribution, densities and carapace width of juvenile T. tridenta-
tus along 6 transects at an intertidal zone of Beibu Gulf, South China Sea, during the summer of 2008.
The environmental parameters measured (water temperature, salinity and dissolved oxygen) did not
vary significantly among transects. However, the abundance and size distribution of juvenile T. tri-
dentatus was statistically different between transects. Juvenile T. tridentatus prefer to move further
from their natal area as they grow: therefore, smaller juveniles tend to be distributed closer to the
shore, while larger juveniles are located further offshore.
KEY WORDS: Chinese horseshoe crab · Tachypleus tridentatus · Distribution · Abundance ·
Size variation · Intertidal zone · Beibu Gulf
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