1 23 Conservation Genetics ISSN 1566-0621 Volume 15 Number 4 Conserv Genet (2014) 15:757-770 DOI 10.1007/s10592-014-0576-1 Range-wide comparisons of northern leatherside chub populations reveal historical and contemporary patterns of genetic variation Jason R. Blakney, Janet L. Loxterman & Ernest R. Keeley