Research Article
Resistance Determinants and Their Association with
Different Transposons in the Antibiotic-Resistant
Streptococcus pneumoniae
Izabela Korona-Glowniak, Radoslaw Siwiec, and Anna Malm
Department of Pharmaceutical Microbiology, Medical University of Lublin, Chodzki 1, 20-093 Lublin, Poland
Correspondence should be addressed to Izabela Korona-Glowniak; iza.glowniak@umlub.pl
Received 22 December 2014; Revised 16 March 2015; Accepted 16 March 2015
Academic Editor: Henny C. van der Mei
Copyright © 2015 Izabela Korona-Glowniak et al. his is an open access article distributed under the Creative Commons
Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is
properly cited.
Multiple resistance of Streptococcus pneumoniae is generally associated with their unique recombination-mediated genetic plasticity
and possessing the mobile genetic elements. he aim of our study was to detect antibiotic resistance determinants and conjugative
transposons in 138 antibiotic-resistant pneumococcal strains isolated from nasopharynx of healthy young children from Lublin,
Poland. hese strains resistant to tetracycline and/or to chloramphenicol/erythromycin/clindamycin were tested by PCR using the
speciic genes as markers. he presence of Tn916 family transposons, carrying tet(M) and int/xisTn916, was observed in all of the
tested strains. Tn916 was detected in 16 strains resistant only to tetracycline. Tn6002 and Tn3872-related element were found among
99 erm(B)-carrying strains (83.8% and 3.0%, resp.). Eight strains harbouring mef (E) and erm(B) genes were detected, suggesting
the presence of Tn2010 and Tn2017 transposons. Among 101 chloramphenicol-resistant strains, two variants of Tn5252-related
transposon were distinguished depending on the presence of int/xis5252 genes speciic for cat gene-containing Tn5252 (75.2% of
strains) or
Sp23FST81
gene, speciic for cat-containing ICESp23FST81 element (24.8% of strains). In 6 strains Tn916-like and Tn5252-
like elements formed a Tn5253-like structure. Besides clonal dissemination of resistant strains of pneumococci in the population,
horizontal transfer of conjugative transposons is an important factor of the high prevalence of antibiotic resistance.
1. Introduction
Antimicrobial resistance among Streptococcus pneumoniae
has spread worldwide and it causes higher risk of treat-
ment failure in pneumococcal infections. S. pneumoniae is
an important human pathogen associated with respiratory
tract infections, bacteriemia, and meningitis. Primarily as a
commensal, the pneumococcus colonizes the nasopharynx of
10–20% of healthy adults and 40–77% of healthy children [1].
Nasopharyngeal carriage in healthy children is a major factor
in the horizontal transmission of pneumococcal strains, espe-
cially between children attending day care centers (DCCs)
or to other family members, and may be also the source
of infection in other individuals. Moreover, pneumococcal
nasopharyngeal isolates relect the strains currently circulat-
ing in the community [1, 2].
Multiple resistance of pneumococci especially resistance
to macrolides and tetracyclines as well as to chloramphenicol
is generally associated with their unique recombination-
mediated genetic plasticity and possessing the mobile genetic
elements [3, 4]. Two major mechanisms of macrolide resis-
tance in S. pneumoniae are noted. he irst one is target site
modiication by a ribosome methylase, encoded by erm(B)
gene and related to high-level resistance to macrolide, lin-
cosamides, and streptogramin B (MLS
B
phenotype). Erm(B)
resistance can be expressed by pneumococci either constitu-
tively (cMLS
B
phenotype) or inducibly (iMLS
B
phenotype)
[5]. he majority of macrolide-resistant S. pneumoniae strains
are also resistant to tetracycline. his association is due
to the insertion of erm(B) into conjugative and composite
transposons of the Tn916 family that harbours tet(M) gene,
encoding ribosome protection proteins. Members of this
Hindawi Publishing Corporation
BioMed Research International
Volume 2015, Article ID 836496, 6 pages
http://dx.doi.org/10.1155/2015/836496