J. Gen. Virol. 2013. 94, 2515-2523. DOI:10.1099/vir.0.055673-0 1 A novel spliced gene in alcelaphine herpesvirus 1 encodes a glycoprotein which is secreted in vitro George C. Russell 1# , Helen Todd 1 , David Deane 1 , Ann Percival 1 , Mark P. Dagleish 1 , David M. Haig 2 and James P. Stewart 3# 1 Moredun Research Institute, Pentlands Science Park, Penicuik, EH26 0PZ, UK 2 School of Veterinary Medicine and Science, Nottingham University, Sutton Bonington, Leicestershire LE12 5RD, UK 3 Department of Infection Biology, Institute of Infection and Global Health, University of Liverpool, Liverpool Science Park IC2, L3 5RF, UK Corresponding authors: George C. Russell, george.russell@moredun.ac.uk ; Tel: +44 131 445 6149; Fax: +44 1313 445 6111 James P. Stewart , j.p.stewart@liv.ac.uk ; Tel: +44 151 795 0221 Footnote: Nucleotide sequences determined in this work have been submitted to the European Nucleotide Archive and have been assigned accession numbers as follows: cDNA clone 001-D10, HE855448; cDNA clone 01-B01, HE855449; cDNA clone 002-H06, HE855450; A9.5*01 allele, HE855451; A9.5*02 allele, HE855452; synthetic expression cassette MCS-HA, FN553440. This is not the version of record of this article. This is an author accepted manuscript (AAM) that has been accepted for publication in [insert name of journal] that has not been copy-edited, typeset or proofed. The Society for General Microbiology (SGM) does not permit the posting of AAMs for commercial use or systematic distribution. SGM disclaims any responsibility or liability for errors or omissions in this version of the manuscript or in any version derived from it by any other parties. The final version is available at DOI:10.1099/vir.0.055673-0 [2013]. 1