24 Herpetological Review 38(1), 2007 Printing Co., Auburn, Alabama. 347 pp. PERRILL, S. A., AND R. E. DANIEL. 1983. Multiple egg clutches in Hyla regilla, H. cinerea, and H. gratiosa. Copeia 1983:513–516. SCARLATA, J. K., AND C. G. MURPHY. 2003. Timing of oviposition by fe- male barking treefrogs (Hyla gratiosa). J. Herpetol. 37:580–582. TRAVIS, J. 1983. Variation in development patterns of larval anurans in temporary ponds. I. Persistent variation within a Hyla gratiosa popu- lation. Evolution 37:496–512. Herpetological Review, 2007, 38(1), 24–30. © 2007 by Society for the Study of Amphibians and Reptiles Herpetofauna of Mount Roraima, Guiana Shield Region, Northeastern South America ROSS D. MACCULLOCH and AMY LATHROP Centre for Biodiversity and Conservation Biology, Royal Ontario Museum 100 Queen’s Park, Toronto, Ontario M5S 2C6, Canada e-mail: rossm@rom.on.ca; amyl@rom.on.ca ROBERT P. REYNOLDS USGS Patuxent Wildlife Research Center, National Museum of Natural History MRC 111, P.O. Box 37012, Washington, D.C. 20013-7012, USA e-mail: reynolds@si.edu J. CELSA SEÑARIS Museo de Historia Natural La Salle, Apartado 1930 Caracas 1010-A, Venezuela e-mail: josefa.senaris@fundacionlasalle.org.ve and GREGORY E. SCHNEIDER Museum of Zoology, University of Michigan Ann Arbor, Michigan 48109-1079, USA e-mail: ges@umich.edu The Guiana region of northeastern South America is an area of high biodiversity, and the varied habitats on the tepuis of the re- gion support a significant portion of this diversity. The zoogeog- raphy of the pantepui region has been a recent topic of interest, with several sources of published data (Duellman 1999; Gorzula and Señaris 1999; Hollowell and Reynolds 2005a; Hoogmoed 1979a; McDiarmid and Donnelly 2005). The herpetofaunal communities of several tepuis have been de- scribed (Donnelly and Myers 1991; Gorzula 1992; McDiarmid and Paolillo 1988; Myers 1997; Myers and Donnelly 1996, 1997, 2001). Mount Roraima, the most famous of these table mountains, has been explored extensively (McDiarmid and Donnelly 2005). Although numerous specimens have been collected on Roraima, these collections have never been summarized or analyzed. Roraima (05º12'N, 060º44'W) is one of the highest points in northeastern South America, and it marks the boundary between Guyana, Venezuela, and Brazil. The summit plateau, from 2600 to 2810 m elevation, is some 34 km 2 in area. The plateau is known for its varied rock formations, and for its very sparse vegetation. The summit is at the top of steep walls which extend from below 2000 m up to 2600–2700 m; these walls are occasionally broken by more gradual slopes. At the bottom of the vertical walls, below about 2000 m, the tepui is surrounded by forested slopes. The for- est is most extensive below 1500 m, with only a narrow band en- circling the mountain above this elevation. Descriptions of the physical features and vegetation of Roraima are in Huber (1995a, b). Herpetofaunas on tepui summits often differ in species compo- sition from faunas on tepui slopes. Summit faunas are often more depauperate than slope faunas (Myers and Donnelly 2001). Both elevation and microhabitat can affect species distributions, and many species occur over a range of elevations (McDiarmid and Donnelly 2005). It is therefore worthwhile to have a compilation of slope and summit species, for comparison with faunas on other mountains. The first aim of this paper is to compile a list of the known herpetofauna of Roraima, from both summit and slopes. The sec- ond aim is to compare the Roraima herpetofauna with those of other tepuis in the Guiana Shield (following Hollowell and Reynolds 2005b, p.1); this consists of 1) comparison using crite- ria developed by McDiarmid and Donnelly (2005), and 2) the hy- potheses of tepui zoogeography of Myers and Donnelly (2001), who enumerated five general points about the composition of tepui herpetofaunas. Such comparisons can provide valuable insight about the zoogeography of the region. Methods.—Information was taken from museum records or pub- lished literature. Amphibian taxonomy follows Faivovich et al. (2005), Frost et al. (2006) and Grant et al. (2006); reptile tax- onomy follows Avila-Pires (2005). Institutional abbreviations fol- low Leviton et al. (1985), with the following additions: CSBD – Centre for the Study of Biological Diversity, University of Guyana, Georgetown, Guyana; MHNLS – Museo de Historia Natural La Salle, Caracas, Venezuela (formerly SCNLS); ULABG – Laboratorio de Biogeografia, Universidad de Los Andes, Mérida, Venezuela. The aims of this study require that two points be defined. First, what are the boundaries of Roraima? Second, what are highland species? These questions were resolved as follows: Roraima: We defined the boundaries of Roraima as that part of the mountain which is above 1500 m. Although some studies of the Guiana Shield region have used 1000 m as a minimum eleva- tion (e.g., Hoogmoed 1979a), the use of this criterion at Roraima would necessitate the inclusion of the extensive surrounding up- lands, and would increase the boundaries of Roraima to an unreal- istic extent. We therefore adopted the 1500 m criterion, as have other studies of the Guiana fauna (Gorzula and Señaris 1999; McDiarmid and Donnelly 2005). Because of the potentially great difference between faunas on tepui summits and faunas on forested tepui slopes, we have indi- cated the location from which each species was recorded, in order to determine whether each is part of the summit or slope faunal assemblages. The habitat on the slopes of Roraima is varied, rang- ing from steep rocky walls to more gently sloping forested areas (Huber 1995a, b). Highland Species: Highland species are those which typically occur above 1500 m; any such species is here considered a high- land species, although it may have been occasionally collected below 1500 m. McDiarmid and Donnelly (2005) followed a simi- lar procedure. Although a highland species may occasionally oc- cur at elevations below 1500 m, it will not be widespread below that elevation. Table 1 contains all species reported from above 1500 m, plus several highland species collected slightly below