African Journal of Aquatic Science 2009, 34(3): 255–259
Printed in South Africa — All rights reserved
Copyright © NISC (Pty) Ltd
AFRICAN JOURNAL OF
AQUATIC SCIENCE
ISSN 1608–5914 EISSN 1727–9364
doi: 10.2989/AJAS.2009.34.3.7.983
African Journal of Aquatic Science is co-published by NISC (Pty) Ltd and Taylor & Francis
Blood lactate levels as a biomarker for angling-induced stress in tigerfish
Hydrocynus vittatus from the Okavango Delta, Botswana
NJ Smit
1
*, G Howatson
1,2
and R Greenfield
1
1
Centre for Aquatic Research, Department of Zoology, University of Johannesburg, Auckland Park, PO Box 524,
Johannesburg 2006, South Africa
2
School of Psychology and Sport Sciences, Northumbria University, Newcastle upon Tyne, NE1 8ST, UK
* Corresponding author, e-mail: nicos@uj.ac.za
Received 12 May 2009, accepted 18 August 2009
Introduction
The tigerfish Hydrocynus vittatus is not only the most
sought-after freshwater angling fish in Africa, but is one
of the most important predatory fishes in African waters
(Winemiller and Kelso-Winemiller 1994, Skelton 2001,
Økland et al. 2005). Although abundant in certain areas
throughout Africa, its numbers have declined in many rivers
due to water extraction, pollution, obstructions such as dams
and weirs, and fishing pressure (Steyn et al. 1996, Skelton
2001). This has been recognised specifically in South Africa
by the Department of Environmental Affairs and Tourism
who recently included tigerfish on the protected species list,
together with such marine icons as the great white shark
Carcharodon carcharius and the once thought to be extinct
coelacanth Latimeria chalumnae (DEAT 2007). Despite
the high profile of the tigerfish as a protected ecologically
and economically important species, only limited published
information on its biology in specific populations is available.
These include data on growth, feeding and population
dynamics in Lake Kariba (Matthes 1968, Kenmuir 1973);
movement patterns and ecology in the upper Zambezi River
and Zambezi floodplain (Winemiller and Kelso-Winemiller
1994, Økland et al. 2005); allozyme variation in and
between upper Zambezi River (Namibia) and Olifants River
(South Africa) populations (Kotzé et al. 1998); and accumu-
lation of selected metals in the tissue and organs of the
Olifants River tigerfish (du Preez and Steyn 1992) and
of mercury in the Lake Kariba tigerfish (Mhlanga 2000).
To date, the majority of research on tigerfish has been on
populations from the Zambezi and Olifants River systems
with no biological information available on the populations
found in the Okavango Delta, Botswana.
The Okavango Delta, specifically the panhandle region
with deeper, faster flowing water, is one of the most popular
areas in Africa for anglers to catch tigerfish (for the specific
region, see van der Bank and Smit 2007, Figure 1). Many
tourist lodges and camps in this area specifically cater for
recreational anglers and are the source for valuable income
for this region. While some anglers keep a portion of the
fish captured (up to five per day are permitted), many fish
are captured and immediately released. The game fishing
industry further encourages anglers voluntarily to release
fish as a way to expand recreational fishing and therefore
catch-and-release angling is growing, as a proportion of
total fishing, in this region. The ultimate success of catch-
and-release angling, however, depends on ensuring high
release survival rates by minimising injury and mortality
(Bartholomew and Bohnsack 2005). Despite the importance
of this principle, research examining the physiological
response to angling and possible sublethal impacts thereof
on freshwater game fish is only available for a few popular
North American and European species, i.e. Atlantic salmon,
rainbow trout and largemouth bass (Brobbel et al. 1996,
Cooke et al. 2002, DuBois and Dubielzig 2004), and is
completely lacking for any of Africa’s freshwater game
Although critical in catch-and-release angling, no data are available on angling-induced stress in African gamefish. Blood lactate
levels were used as a biomarker for angling-induced metabolic stress in tigerfish caught by angling in the Okavango Delta,
Botswana. Blood was drawn and analysed for blood lactate from 66 anaesthetised fish. The landing time, handling time, body
mass and total length were recorded prior to reviving and keeping the fish in aerated water for recovery before release. A strong
positive relationship (r
2
= 0.607) was found between landing time and body mass, as well as significant elevations in blood lactate
concentrations following rod-and-line angling, regardless of angling time. These data suggest that longer angling time significantly
increases physiological stress, which may have an impact on breading success and mortality in tigerfish.
Keywords: catch-and-release angling, fish, physiology, recreational fisheries management, stress