Karyotypic analyses and morphological comments on the endemic
and endangered Brazilian painted tree rat Callistomys pictus
(Rodentia, Echimyidae)
Karen Ventura
1
, Gilson Evaristo Iack Ximenes
2
, Renata Pardini
3
, Marcos A. Nóbrega de Sousa
4
,
Yatiyo Yonenaga-Yassuda
1
and Maria José de J. Silva
5
1
Departamento de Genética e Biologia Evolutiva, Instituto de Biociências, Universidade de São Paulo,
São Paulo, SP, Brazil.
2
Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz, Ilhéus, BA, Brazil.
3
Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, SP, Brazil.
4
Centro de Ciências Biológicas e Sociais Aplicadas, Universidade Estadual da Paraíba, João Pessoa,
PB, Brazil.
5
Laboratório Especial de Ecologia e Evolução, Instituto Butantan, São Paulo, SP, Brazil.
Abstract
The genus Callistomys belongs to the rodent family Echimyidae, subfamily Echimyinae, and its only living represen-
tative is Callistomys pictus, a rare and vulnerable endemic species of the state of Bahia, Brazil. Callistomys has been
previously classified as Nelomys, Loncheres, Isothrix and Echimys. In this paper we present the karyotype of
Callistomys pictus, including CBG and GTG-banding patterns and silver staining of the nucleolus organizer regions
(Ag-NORs). Comments on Callistomys pictus morphological traits and a compilation of Echimyinae chromosomal
data are also included. Our analyses revealed that Callistomys can be recognized both by its distintinctive morphol-
ogy and by its karyotype.
Key words: Callistomys, karyotype, banding patterns, Echimyinae.
Received: September 28, 2007; Accepted: March 17, 2008.
The Neotropical spiny rats of the family Echimyidae
comprise approximately 85 recognized living species
grouped in 21 genera (modified from McKenna and Bell,
1997; Woods and Kilpatrick, 2005; Emmons, 2005; Iack-
Ximenes et al., 2005). Echimyids have a wide distribution,
ranging from Southern Paraguay to Southern Nicaragua
(Anderson and Jones, 1967; Honacki et al., 1982; Harten-
berger, 1985). They are the most diverse of all living hystri-
cognath rodents and the evolutionary relationships among
genera are still unclear due to the paucity of data regarding
their ecology, diversity and systematics (Leite and Patton,
2002; Woods and Kilpatrick, 2005).
Woods and Kilpatrick (2005) recognized four Echi-
myidae subfamilies: (1) the extinct Heteropsomyinae, en-
demic to West India; (2) Dactylomyinae, including the
arboreal bamboo rats from the genera Dactylomys,
Kannabateomys and Olallamys; (3) Echimyinae, including
the arboreal genera Callistomys, Diplomys, Echimys,
Phyllomys, Isothrix and Makalata; and (4) Eumysopinae,
including the semi-fossorial, arboreal and terrestrial genera
Carterodon, Clyomys, Euryzygomatomys, Lonchothrix,
Mesomys, Hoplomys, Proechimys, Trinomys and
Thrichomys. Since then, several new genera of Echimyinae
have been erected: Pattonomys to include Nelomys
semivillosus Geoffroy, 1838 and allied species,
Santamartamys to include Isothrix rufodorsalis Allen,
1899 (Emmons, 2005) and Toromys to include Loncheres
grandis Wagner 1845 (Iack-Ximenes et al., 2005).
Karyotypes of Echimyinae are known only for
Pattonomys semivillosus and for some species of
Phyllomys, Makalata and Isothrix and their chromosome
numbers ranged from 2n = 22 in Isothrix pagurus to 2n = 96
in Phyllomys medius (Table 1).
The painted tree rat or cocoa rat Callistomys pictus
(Pictet, 1843) is a soft-furred echimyid found in the coast of
the state of Bahia, Northeastern Brazil. It was originally in-
cluded in the genus Nelomys as a junior synonym of
Echimys. It was afterwards alternatively classified as
Genetics and Molecular Biology, 31, 3, 697-703 (2008)
Copyright © 2008, Sociedade Brasileira de Genética. Printed in Brazil
www.sbg.org.br
Send correspondence to Maria José de Jesus Silva. Laboratório
Especial de Ecologia e Evolução, Instituto Butantan, Av. Dr. Vital
Brazil 1500, 05503-900 São Paulo, SP, Brazil. E-mail: mariajo@
butantan.gov.br or mariajo@usp.br.
Short Communication