Brief report
Occurrence of a monophasic strain of Salmonella
group B isolated from cetaceans in England and Wales
between 1990 and 2002
Carlos A. Valderrama Vasquez,
1,2
*
Shaheed K. Macgregor,
1
J. Marcus Rowcliffe
1
and
Paul D. Jepson
1
1
Institute of Zoology, Zoological Society of London,
Regent’s Park, London NW1 4RY, UK.
2
Fundación Vida Silvestre Neotropical, Carrera 65b no.
10
a
-05, Bogotá DC, Colombia.
Summary
Between September 1990 and December 2002, 511
cetacean carcasses stranded or caught in commer-
cial fisheries in England and Wales were examined
post mortem. Salmonella group B was isolated from
60 of 279 (21.51%) harbour porpoises (Phocoena pho-
coena), predominantly from lung tissue. Forty-three
of the Salmonella group B isolates were subsequently
serotyped and all found to have the antigenic struc-
ture O4,12:a:-. The annual proportion of harbour
porpoises testing positive for Salmonella O4,12:a:-
increased significantly from 6% in the early 1990s to
27% after 1999. The cause(s) of the increasing preva-
lence of Salmonella O4,12:a:- are not known, but may
reflect natural variation in the epidemiological cycle
of Salmonella O4,12:a:- in harbour porpoises.
The probability of isolating Salmonella O4,12:a:-
from harbour porpoises increased with age, suggest-
ing that the mode of transmission is principally
horizontal. There appeared to be a weak degree of
seasonality in the probability of isolating Salmonella
O4,12:a:- with a low proportion of cases in the months
of April and May. Based on pathological findings from
infected carcasses, Salmonella O4,12:a:- may be part
of the normal commensal flora of the lungs of ceta-
ceans with the potential to act as an opportunistic
pathogen.
Introduction
The genus Salmonella is a well-recognized pathogen
of people, livestock, wild animals, birds, reptiles and
even insects. Microscopically, Salmonella spp. are small,
straight, Gram-negative rods (0.7–1.5 ¥ 2.0–5.0 mm).
Most are motile but non-motile forms also occur (Williams
and Barker, 2001). Bacterial diseases are a significant
cause of death in marine mammals, either as a primary
cause or secondary to viral or helminth infections
(Moeller., 1997; Dunn et al., 2001).
In cetaceans, Salmonella spp. bacteria are common
pathogens and may cause severe necrotizing enteritis
and/or septicaemia in affected animals (Kirkwood et al.,
1997; Moeller, 1997; Jepson et al., 2000; Tryland, 2000;
Jepson, 2005). Clinically, animals have been reported to
show severe enteritis with necrosis and/or haemorrhage
and septicaemic cases usually die without showing
clinical signs. In some cases, septicaemia can lead to
complications such as bronchopneumonia, necrotizing
hepatitis, meningoencephalitis and abscessation
(Moeller, 1997). Isolates of Salmonella spp. have been
reported from killer whales (Orcinus orca), bottlenose dol-
phins (Tursiops truncatus) and pilot whales (Globicephala
melas) (Tryland, 2000), and Salmonella DT12 has been
reported from a harbour porpoise (Foster et al., 1999).
Furthermore, there have been reports of large outbreaks
of salmonellosis in humans who have eaten whale meat
or blubber, with clinical signs including fever, chills,
nausea, vomiting, cramps and diarrhoea (Tryland, 2000).
Although the primary site of infection of Salmonella spp.
is the gastrointestinal tract (Williams and Barker, 2001), a
monophasic group B Salmonella with the antigenic struc-
ture O4,12:a:- has been isolated from harbour porpoises
(Phocoena phocoena) where the lung is the tissue most
frequently infected (Foster et al., 1999). It has been sug-
gested that this particular serotype of Salmonella is host-
adapted and opportunistic because it has been isolated
from pneumonic and non-pneumonic lungs, including
from the lungs of healthy harbour porpoises that were
accidentally caught in fishing nets or killed by bottlenose
Received 25 April, 2007; accepted 9 March, 2008. *For
correspondence. E-mail carlos.valderrama@fvsn.org; Tel. (+57)
3114668425; Fax (+57) 12611942.
Environmental Microbiology (2008) 10(9), 2462–2468 doi:10.1111/j.1462-2920.2008.01651.x
© 2008 The Authors
Journal compilation © 2008 Society for Applied Microbiology and Blackwell Publishing Ltd