ISSN 0031-0301, Paleontological Journal, 2015, Vol. 49, No. 4, pp. 413–423. © Pleiades Publishing, Ltd., 2015.
Original Russian Text © V.V. Bulanov, A.G. Sennikov, 2015, published in Paleontologicheskii Zhurnal, 2015, No. 4, pp. 79–90.
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INTRODUCTION
Reliable remains of the most ancient gliding rep-
tiles, Permian Weigeltisauridae, are presently known
from Madagascar, Germany, Great Britain, and Rus-
sia. The first taxon of this group was formally described
by Piveteau (1926) under the name Coelurosauravus
elivensis from Gondwana, referred to coelurosaurs,
and regarded as a probable ancestor of Compsognathus.
At present, the collection of C. elivensis is represented
by imprints of three skeletons. Since the bones are
completely weathered out, the morphology of this spe-
cies is usually described based on the plastic replicas of
imprints. The material from Madagascar is of great
importance for a better understanding of the weigelti-
saurid postcranium, including the axial region, limbs,
girdles, and framework structures of the patagium
(Carroll, 1978). The skull of C. elivensis, the morpho-
logical study of which is particularly important for
revision of the generic composition of the family,
could have been reconstructed only partially and the
present study is intended to solve this task.
Until now, in the study of cranial morphology of
weigeltisaurids, attention was paid to the material
from Europe. The first known European member of
the family was described under the name Palaeo-
chamaeleo jaekeli from the Kupferschiefer of Ger-
many (Weigelt, 1930). Since this generic name turned
out to be preoccupied, it was subsequently replaced by
Weigeltisaurus (Kuhn, 1939). For a long time, the skull
of the holotype of W. jaekeli (specimen SSWG,
no. 113/7), which is much more completely preserved
than the specimens from Madagascar, was the basis for
reconstruction of the weigeltisaurid skull, discussion
of homology of its elements, and taxonomic position
of the group.
The last revision of the type collection of C. eliven-
sis and W. jaekeli resulted in the rejection of the genus
Weigeltisaurus (Evans and Haubold, 1987). All West
European specimens found later, such as an aberrant
specimen from Mansfeld (Schaumberg et al., 2007),
“the specimen from Eppleton” (Pettigrew, 1979),
which is only known from the postcranium, and a
remarkable skeleton from Karlsruhe (Frey et al., 1997)
were assigned to the genus Coelurosauravus. In the
present study, we accept the validity of the genus
Weigeltisaurus, taking into account that it differs from
Coelurosauravus in the structure of cervical vertebrae
and some bones of the skull roof, as follows from the
material described below and available data on the
morphology of W. jaekeli (Weigelt, 1930; Evans and
Haubold, 1987).
Comparative material includes East European
weigeltisaurids from localities of the Kul’chumovo
section in the Orenburg Region (Russia). Our sample
includes mostly isolated bones belonging to individu-
als varying in age and referred to two species of the
genus Rautiania, R. alexandri and R. minichi (Bulanov
and Sennikov, 2006), the differences between which
are observed in the structure of parietals and maxillae.
The holotypes of these species are the parietals, which
New Data on the Morphology of the Late Permian Gliding Reptile
Coelurosauravus elivensis Piveteau
V. V. Bulanov
a
and A. G. Sennikov
a, b
a
Borissiak Paleontological Institute, Russian Academy of Sciences,
Profsoyuznaya ul. 123, Moscow, 117997 Russia
b
Kazan Federal University, Kremlevskaya ul. 18, Kazan, 420008 Russia
e-mail: bulanov@paleo.ru, sennikov@paleo.ru
Received May 14, 2014
Abstract—Based on the material from the type locality (Eliva Mountain, Madagascar), the cranial morphol-
ogy of the Late Permian gliding reptile Coelurosauravus elivensis Piveteau, 1926 (Weigeltisauridae) is
redescribed. The diagnosis of the genus Coelurosauravus is emended based on comparisons with East Euro-
pean weigeltisaurids of the genus Rautiania. It is shown that C. elivensis and all weigeltisaurids lack the post-
frontals and supratemporals and have a small preorbital fenestra between the maxilla and nasal, partially
reduced lacrimal, and large nasolacrimal duct piercing the prefrontal. The poor development of ornamental
outgrowths on the parietal and in the ventral part of the squamosal as well as the small size of studied individ-
uals of C. elivensis are caused by their juvenile age.
Keywords: gliding reptiles, cranial morphology, Late Permian, Madagascar
DOI: 10.1134/S0031030115040048