Partial Purification and Characterisation of Polyphenol Oxidase from Hibiscus
rosa–sinensis L
Ikram Madani
Faculty of Science
University of Khartoum
Khartoum, Sudan
Ikramahmed3@yahoo.com
Pat M. Lee
Materials Technology Program
Universiti Teknologi MARA
Shah Alam, Selangor, Malaysia
patmlee.1@gmail.com
Lee Kong Hung
Department of Bitechnology,
Malaysia University of Science and
Technology
Petaling Jaya, Selangor, Malaysia.
konghunglee@hotmail.com
Abstract— Polyphenol oxidase (PPO) from petals of Hibiscus
rosa–sinensis L. (Chinese Rosa) was extracted and partially
purified by ammonium sulphate precipitation. Its physical and
chemical properties such as optimal pH and temperature,
substrate specificity, and effects of inhibitors and activators on
the enzymatic activity were investigated. The total protein
contents of Hibiscus rosa–sinensis petals crude extract and
partially purified extract were found to be 425 μg and 122 μg
/100 g fresh petals respectively. The enzyme activity and its
kinetic parameters, (K
m
and V
max
) were determined using 4-
methylcatechol and catechol as substrates. The optimum pH
and temperature for Hibiscus PPO activities were 6.0 and
45 °C respectively. Ascorbic acid and
ethylenediaminetetraacetic acid sodium salt were found to be
potent inhibitors and copper sulphate was found to be
activator for Hibiscus PPO.
catechol, 4-methyl catechol , K
m
and V
max
I. INTRODUCTION
Polyphenol oxidase (PPO, EC 1.14.18.1) is a common
copper containing enzyme which is widely distributed in
nature. In the presence of oxygen it catalyses two reactions:
the hydroxylation of monophenols to o–diphenols and the
oxidation of o–diphenols to o–quinones. This enzyme plays
an important role in many plant metabolic processes. The
oxidation of phenol compounds by PPO are polymerized to
form dark–coloured compounds responsible for browning in
fruits and vegetables. This is regarded as an undesirable
sensory attribute of the produce and decreases its commercial
and nutritive values. For this reason PPO has received
increasing attention [1]. PPO enzyme has been isolated and
characterized from produce of many commercially important
plants and fruits such as lettuce [2], aubergine [3], Ferula sp.
[4], strawberry [5], and cocoa beans [6].
The present investigation was designed to extract PPO
from petals of Hibiscus rosa–sinensis (Chinese Rose) which
is the national flower of Malaysia. The enzyme was partially
purified from the aqueous extract of flower petals, and its
substrate specificity, kinetic parameters, optimum conditions
of pH and temperature, and effects of inhibitor and metallic
compounds on the enzymatic activity were evaluated. The
results of this study would provide an understanding of the
browning of the hibiscus flower and means of prolonging the
shelf flower.
II. MATERIALS AND METHODS
All chemicals used were of analytical grade.
Polyvinylpyrrolidone from Acros Organics; ascorbic acid
and copper sulphate from Fisher Scientific; monosodium
phosphate from Fluka Biochemical; disodium phosphate and
iron III chloride from Sigma Chemical Co; 4-methylcatechol
and catechol from Merk–Schuchardt CHG; EDTA from
Progma.
A. Plant Materials
Two hundred grams of fresh red flowers of Hibiscus
rosa–sinensis, grown in Kuala Lumpur, Malaysia, were
hand-picked from the plant and left to dry at room
temperature.
B. Enzyme Extraction and Partial Purification
Ten grams of dried petals were homogenized in 80 mL of
0.1M phosphate buffer (pH 6.8) containing 10 mM ascorbic
acid and 0.5% polyvinylpyrrolidone (PVP40) with the aid of
a magnetic stirrer for 1h. The crude extract samples were
centrifuged at 32000 g for 20 min at 4ºC. Solid ammonium
sulphate (NH
4
)
2
SO
4
was added to the supernatant to obtain
80% (NH
4
)
2
SO
4
saturation. After 1 h, the precipitated
proteins were separated by centrifugation at 32000 g for 30
min. The precipitate was re–dissolved in a small volume of
distilled water and dialyzed at 4ºC against distilled water for
24 h with 4 changes of the water during dialysis. The
dialyzed sample was lyophilized and this constitutes the
partial purified PPO extract and was used as the Hibiscus
PPO enzyme source.
C. Determination of Hibiscus PPO Activity
Hibiscus–PPO activity was determined by measuring the
absorbance at 420 nm using a spectrophotometer
(Pharmatech, Model UV–1700). To determine the best
concentration of enzyme preparation corresponding to the
highest enzyme activity, the activity was assayed in 3 mL of
reaction mixture consisting of 0.5 mL substrate (0.02 M 4–
methylcatechol and 0.02 M catechol separately) and different
concentrations (0.025-0.5 mL) of the enzyme preparation
(1mg/mL). This mixture was topped-up to 3.0 mL with the
phosphate buffer (pH 6.8) in a 1 cm light path quartz cuvette.
The blank consisted of 3.0 mL 0.1 M phosphate buffer (pH
6.8). Two controls were prepared: the cuvette of the first
110
Keywords Polyphenol oxidase, Hibiscus rosa–sinensis; -
2011 2nd International Conference on Biotechnology and Food Science
IPCBEE vol.7 (2011) © (2011) IACSIT Press, Singapore