Development and validation of a 3ABC antibody ELISA in Australia
for foot and mouth disease
A Colling,
a
* C Morrissy,
a
J Barr,
a
G Meehan,
a
L Wright,
a
W Goff,
a
LJ Gleeson,
a
B van der Heide,
a
S Riddell,
a
M Yu,
a
D Eagles,
a
R Lunt,
a
S Khounsy,
b
Ngo Than Long,
c
Pham Phong Vu,
c
Nguyen Than Phuong,
c
Nguyen Tung,
d
W Linchongsubongkoch,
e
J Hammond,
f
M Johnson,
f
WO Johnson,
g
H Unger,
h
P Daniels
a
and JR Crowther
h
Objective To measure the diagnostic performance of an
Australian-developed ELISA for the detection of antibodies against
the non-structural proteins (NSP) 3ABC of the foot and mouth
disease (FMD) virus.
Design Test development and validation study.
Methods The diagnostic specificity was determined using 2535
sera from naïve animals and 1112 sera from vaccinated animals.
Diagnostic sensitivity was calculated from the data for 995 sera
from experimentally and field-infected animals from FMD-endemic
countries in South East Asia. A commercial ELISA detecting antibod-
ies against FMD virus NSP was used as the reference test to establish
relative sensitivity and specificity. Bayesian latent class analysis was
performed to corroborate results. The diagnostic window and rate
of detection were determined at different times using sera from
cattle, sheep and pigs before and after infection, and after vaccina-
tion and subsequent infection. Repeatability and reproducibility
data were established.
Results At 35% test cut-off, the 3ABC ELISA had an overall diag-
nostic sensitivity of 91.5% and diagnostic specificity of 96.4%. The
diagnostic sensitivity in vaccinated and subsequently infected
cattle was 68.4% and diagnostic specificity in vaccinated cattle was
98.0%.
Conclusions The 3ABC ELISA identified field and experimentally
infected animals, as well as vaccinated and subsequently infected
animals. Diagnostic sensitivity and specificity estimates for other
FMD NSP tests are comparable with the results obtained in this
study. This NSP ELISA was found to be ‘fit for purpose’ as a screening
assay at the herd level to detect viral infection and also to substan-
tiate absence of infection.
Keywords 3ABC ELISA; foot and mouth disease; screening tests
Abbreviations ASe, analytical sensitivity; ASp, analytical specific-
ity; CI, confidence interval; CV, coefficient of variation; dpi, days post
infection; DSe, diagnostic sensitivity; DSp, diagnostic specificity;
FMD, foot and mouth disease; NSP, non-structural protein; PI,
percentage of inhibition; ROC, receiver-operating characteristic; SD,
standard deviation; SPCE, solid-phase competition ELISA
Aust Vet J 2014;92:192–199 doi: 10.1111/avj.12190
F
oot and mouth disease (FMD) must be differentiated from
other vesicular diseases, so rapid laboratory confirmation of a
suspected case is imperative. Direct diagnosis is by virus iso-
lation or demonstration of antigen or nucleic acid in samples of tissue
or fluid. Detection of antibodies to FMD viral non-structural proteins
(NSPs) is an indicator of infection, irrespective of vaccination status,
1
and is particularly useful in countries where vaccination is used or in
mild cases or where epithelial tissue cannot be collected. Most of the
commercial tests for the detection of antibodies to NSPs use recom-
binant expressed NSP 3ABC target antigens.
2
Antibodies against NSP
indicate past or present infection with any of the seven serotypes of
FMD virus, even if animals have been vaccinated. Recent publications
indicate that the sensitivity and specificity of such assays for the
detection of carriers among vaccinated cattle populations are appro-
ximately 90% and 99%, respectively.
1,3-8
The use of NSP-ELISAs to
substantiate freedom from FMD infection after emergency vaccina-
tion of cattle has become an important option in Europe in recent
years.
9
Vaccine purity is an important consideration when using these assays
because trace amounts of NSP in formulations may result in false-
positive reactions in animals that are repeatedly vaccinated.
1,10
Con-
versely, antibodies against NSPs are not detected by some assays
2,11
in
some FMD-vaccinated animals that have been subsequently chal-
lenged with infectious virus and then confirmed to be persistently
infected. Paradoxically, in some vaccinated cattle that later become
infected, there is a delayed antibody response against NSP compared
with the response in unvaccinated-and-infected cattle.
2,5
Some studies
have shown that anti-NSP tests can identify carriers in the absence
of any previous clinical signs when virus is not detectable by reverse
transcriptase polymerase chain reaction, suggesting that anti-NSP
testing, even of individual animals, may detect infections that may
be missed by other tests.
2,12
For these reasons, NSP assays are best
used at the herd level to detect FMD virus circulation in vaccinated
populations.
For FMD-free countries such as Australia, it is important to develop
and validate test platforms based on a differentiating-infected-from-
vaccinated-animals (DIVA) strategy and to be prepared for FMD
incursions by having an independent supply of reagents. The aim of
this study was to assess an Australian-developed 3ABC ELISA for
*Corresponding author.
a
CSIRO Livestock Industries, Australian Animal Health Laboratory, Geelong, Victoria,
Australia; axel.colling@csiro.au
b
Northern Region Sustainable Livelihoods through Livestock Development Project,
Ministry of Agriculture, DLF Regional Office, Luang Prabang, Lao PDR
c
Regional Animal Health Office No. 6, Ho Chi Minh City (RAHO6-HCMC), Vietnam
d
National Centre for Disease Control, Hanoi, Vietnam
e
Foot and Mouth Disease Centre, Department of Livestock Development, Pakchong,
Thailand
f
Institute for Animal Health, Pirbright Laboratory, Pirbright, Woking, Surrey, UK
g
Department of Statistics, University of California at Irvine, USA
h
Joint FAO/IAEA Division, Vienna, Austria
PRODUCTION ANIMALS
PRODUCTION ANIMALS
© 2014 Australian Veterinary Association Australian Veterinary Journal Volume 92, No 6, June 2014 192