744 CEREAL CHEMISTRY
Basic Rheology of Bread Dough with Modified Protein Content
and Glutenin-to-Gliadin Ratios
S. Uthayakumaran,
1–3
M. Newberry,
1,3
M. Keentok,
1,3
F. L. Stoddard,
1,4
and F. Bekes
1,5
ABSTRACT Cereal Chem. 77(6):744–749
The uniaxial elongational and shear rheology of doughs varying in either
the protein content or glutenin-to-gliadin ratio were investigated. Increasing
the protein content at constant glutenin-to-gliadin ratio increased the strain-
hardening properties of the dough, as shown by increasing elongational
rupture viscosity and rupture stress. Glutenin and gliadin had a more com-
plex effect on the elongational properties of the dough. Increased levels
of glutenin increased the rupture viscosity but lowered the rupture strain,
while elevated gliadin levels lowered the rupture viscosity but increased
the rupture strain. These observations provide rheological support for the
widely inferred role of gliadin and glutenin in shaping bread dough rheology,
namely that gliadin contributes the flow properties, and glutenin contri-
butes the elastic or strength properties. The shear and elongational prop-
erties of the doughs were quite different, reflecting the dissimilar natures
of these two types of flow. Increasing protein content lowered the maxi-
mum shear viscosity, while increasing the glutenin-to-gliadin ratio increased
maximum shear viscosity. Strong correlations between the results of
basic and empirical rheology were found. These basic, or fundamental,
rheological measurements confirmed prior empirical studies and supported
baking industry experience, highlighting the potential of basic rheology
for bread and wheat research.
Dough measurements, as determined on equipment such as the
mixograph, farinograph, extensigraph and alveograph, all utilize arbi-
trary units that cannot readily be converted to more useful scientific
dimensions (Levine 1987). Basic rheological instruments are cap-
able of providing the essential, or fundamental, details of the
material’s rheological properties, unlike empirical tests. Rheological
properties can be considered as a continuum between two ideal
states, those of pure elasticity and of pure viscosity. A purely
elastic rubber band does not flow but snaps back to its starting
position, while purely viscous water flows without recovery. Most
materials, including bread dough, demonstrate behavior that is a
combination of both states and basic rheometry can elucidate and
quantify these properties.
Wheat dough is a unique material formed when wheat flour is
mixed with water creating a viscoelastic dough that retains gas
(Walker and Hazelton 1996). The rheological characterization of
wheat flour dough is essential to produce information concerning
the quality of the raw material and the textural characteristics of
the finished product. Dough studies in which the basic principles
of physics have been applied have involved basic rheological
measurements of shear stress in steady shear, creep, stress
relaxation and extension (Bloksma and Bushuk 1988, Janssen et al
1996a, Safari-Ardi and Phan-Thien 1998). These studies have had
the aim of completely characterizing dough and of finding reliable
rheological tests that can differentiate dough types. Strain sweep
experiments and stress-relaxation tests on Australian strong flour
indicated that tests at higher shear strains can differentiate flour
types (Phan-Thien and Safari-Ardi 1998, Safari-Ardi et al 1998).
A study on the effects of starch-protein interaction on rheological
properties has also been performed on two South Australian wheat
cultivars where a synthetic aqueous mixture of dough was used
(Chiruta et al 1997). Khatkar et al (1995) and Janssen et al (1996a)
studied the effect of varying the glutenin-to-gliadin ratio on rheo-
logical property of gluten dough and showed that glutenin contri-
buted to elastic and gliadin to the viscous property of hydrated
gluten. Though it has been shown that basic rheological tests can
differentiate flour samples, very little work of this nature has been
reported so far on the relationship between different protein quan-
tities, different glutenin-to-gliadin ratios, and their functional
properties.
The objective of this study was to use basic rheological tests
(elongational and shear viscometry) to separate the effects of
protein quantity and composition on dough properties.
MATERIALS AND METHODS
Samples
Wheat flours Banks, Hartog, Rosella, and Sunbri were obtained
from BRI Australia Ltd., North Ryde, NSW, for the study. The
high molecular weight glutenin subunit (HMW-GS) composition
of the flours is given in Table I.
The flour components (starch, gluten, glutenin-rich and gliadin-
rich fractions) for enrichment studies were prepared as described
by MacRitchie (1987) and Uthayakumaran et al (1999). The
nitrogen content of the components was determined by the Dumas
total combustion method using an elemental analyzer (CHN-1000,
Leco Inc., St. Joseph, MI). Protein (%) was estimated as N × 5.7.
Altering Protein Content of Flour
at Constant Glutenin-to-Gliadin Ratio
Blends (10.0 g) of each of the base flours, using gluten and
starch isolated from that flour, were prepared as previously des-
cribed (Uthayakumaran et al 1999). Based on the protein content
of each flour, gluten and starch blends of flour and gluten isolated
from it were prepared to have 110, 120, and 130% of the protein
content in the base flour (increasing protein). Formulations contain-
ing 80 and 90% of the protein of the parent flour were prepared by
blending the flour with starch isolated from that flour (to dilute the
protein).
Altering Glutenin-to-Gliadin Ratio
at Constant Protein Content
Gluten, glutenin, or gliadin prepared from the parent flour were
added to the flour to vary the glutenin-to-gliadin ratio while keeping
the protein content constant at 120% of the protein content of the
parent flour.
Measurement of Functional Properties
The amounts of water to be added were calculated from the
protein content and the moisture of the blend using the standard
method (AACC 2000). Blend, water, and salt solution (6.67%
1
Quality Wheat Cooperative Research Centre Ltd., Locked Bag 1345, North
Ryde, NSW 1670, Australia.
2
Corresponding author. Phone: 61 2 9351-7141. Fax: 61 2 93517060. E-mail:
suthay@mech.eng.usyd.edu.au
3
Department of Mechanical and Mechatronic Engineering, The University of
Sydney, NSW 2006, Australia.
4
Plant Breeding Institute, The University of Sydney, NSW 2006, Australia.
5
CSIRO Plant Industry, Grain Quality Research Laboratory, P.O. Box 7, North
Ryde, NSW, 1670, Australia.
Publication no. C-2000-1011-03R.
© 2000 American Association of Cereal Chemists, Inc.