© 2014 Wiley-VCH Verlag GmbH & Co. KGaA, Weinheim 1
1 Introduction
Recent developments of biotransformation processes
have been driven by the increasing demand for sustain-
able, energy-saving, and environmentally friendly syn-
thetic strategies [1, 2]. Although enzymes are biocatalysts
with high efficiency and excellent selectivity, their wide-
spread application in industrial processes is often limited
due to poor reusability and low operational stability [3].
Many immobilization strategies have been developed to
improve biocatalyst properties [4–7]. Maximum retention
of the enzymatic activity has been the primary concern,
whereas less attention has been paid to the biocatalyst
structure and its changes during immobilization and use
under process conditions. This is surprising considering
Research Article
Microscopic monitoring provides information on structure
and properties during biocatalyst immobilization
Sarka Bidmanova
1,2
, Eva Hrdlickova
3
, Josef Jaros
4
, Ladislav Ilkovics
4
, Ales Hampl
2,4
, Jiri Damborsky
1,2,3
and Zbynek Prokop
1,2,3
1
Loschmidt Laboratories, Department of Experimental Biology and Research Centre for Toxic Compounds in the Environment
RECETOX, Faculty of Science, Masaryk University, Brno, Czech Republic
2
International Clinical Research Center, St. Anne’s University Hospital Brno, Czech Republic
3
Enantis Ltd., Brno, Czech Republic
4
Department of Histology and Embryology, Faculty of Medicine, Masaryk University, Brno, Czech Republic
Enzymes have a wide range of applications in different industries owing to their high specificity
and efficiency. Immobilization is often used to improve biocatalyst properties, operational stabil-
ity, and reusability. However, changes in the structure of biocatalysts during immobilization and
under process conditions are still largely uncertain. Here, three microscopy techniques – bright-
field, confocal and electron microscopy – were applied to determine the distribution and structure
of an immobilized biocatalyst. Free enzyme (haloalkane dehalogenase), cross-linked enzyme
aggregates (CLEAs) and CLEAs entrapped in polyvinyl alcohol lenses (lentikats) were used as
model systems. Electron microscopy revealed that sonicated CLEAs underwent morphological
changes that strongly correlated with increased catalytic activity compared to less structured, non-
treated CLEAs. Confocal microscopy confirmed that loading of the biocatalyst was not the only fac-
tor affecting the catalytic activity of the lentikats. Confocal microscopy also showed a significant
reduction in the pore size of lentikats exposed to 25% tetrahydrofuran and 50% dioxane. Narrow
pores appeared to provide protection to CLEAs from the detrimental action of cosolvents, which
significantly correlated with higher activity of CLEAs compared to free enzyme. The results showed
that microscopy can provide valuable information about the structure and properties of a biocat-
alyst during immobilization and under process conditions.
Keywords: Cross-linked enzyme aggregates · Immobilization · Microscopy · Polyvinyl alcohol · Structure
Correspondence: Assoc. Prof. Zbynek Prokop, Loschmidt Laboratories,
Department of Experimental Biology, Faculty of Science, Masaryk
University, Kamenice 5/A13, 625 00 Brno, Czech Republic
E-mail: zbynek@chemi.muni.cz
Additional Correspondence: Prof. Jiri Damborsky, Loschmidt Laboratories,
Department of Experimental Biology, Faculty of Science, Masaryk
University, Kamenice 5/A13, 625 00 Brno, Czech Republic
E-mail: jiri@chemi.muni.cz
Abbreviations: CLEAs, cross-linked enzyme aggregates; lentikats, lens-
shaped particles of polyvinyl alcohol
Biotechnol. J. 2014, 9 DOI 10.1002/biot.201300049
www.biotechnology-journal.com
Biotechnology
Journal
Received 29 JAN 2014
Revised 31 JAN 2014
Accepted 14 MAR 2014
Accepted
article online 17 MAR 2014
Supporting information
available online