SHALLOW LAKES
Reed stands during different water level periods:
physico-chemical properties of the sediment and
growth of Phragmites australis of Lake Balaton
Viktor R. Tóth
Received: 24 March 2015 / Revised: 19 January 2016 / Accepted: 6 February 2016
© Springer International Publishing Switzerland 2016
Abstract Water level fluctuations play a vital role
in regulating macrophytes of shallow lakes. Mor-
phology and growth dynamics of Phragmites
australis, together with physico-chemical parameters
of the sediment, were studied at stable (not degrad-
ing) and die-back (degrading) sites of Lake Balaton
over an 8-year period that included low and average
water levels. Lower water level increased plant
density and green leaf number, positively affecting
photosynthetically available leaf area. Nevertheless,
rhizome carbohydrate content was not influenced by
water level decrease. The physico-chemical parame-
ters of the sediment did not vary greatly, although the
nitrogen and phosphorus content and the midsummer
redox potential of the sediment were higher at the
low-water period. During the transition from average
to low water levels, the sediment shifted from
severely anoxic to poorly oxidised levels, with more
favourable nutrient content, while the amount of
ammonia and sulphides decreased, too. It was shown
that lowering water levels could act on plants via
increased redox potential of the sediment and could
counteract the die-back of Phragmites, suggesting the
effectiveness of water level decrease as a manage-
ment practice to counter reed die-back.
Keywords Redox potential · Morphology ·
Growth dynamics · Carbohydrates ·
Water level changes
Introduction
Water depth is one of the crucial factors that controls
zonation, distribution and progression of Phragmites
australis within lakes (Coops et al., 1996; Vretare
et al., 2001; Engloner & Papp, 2006; To ´th & Szabo ´,
2012). Numerous studies have shown that, due to
specific cytological and biophysical features, com-
mon reed is able to tolerate high and prolonged
inundation (Armstrong et al., 1994; Crawford &
Braendle, 1996; Vartapetian & Jackson, 1997). A
continuous gas space within the plant tissue called
aerenchyma runs down from the aerial to the
underground parts of the plant, channelling air from
leaves to rhizomes and roots. This flux of atmo-
spheric gases is driven by humidity-induced partial
pressure differences between the air and the sub-
stomatal space (Armstrong & Armstrong, 1991).
Without it, the reed could suffer complete or partial
oxygen deprivation due to high microbial oxygen
consumption within the sediment (Brinson et al.,
1981; Crawford & Braendle, 1996). To survive
Guest editors: M. Bekliog ˘lu, M. Meerhoff, T. A. Davidson,
K. A. Ger, K. E. Havens & B. Moss / Shallow Lakes in a Fast
Changing World
V. R. Tóth (&)
Hungarian Academy of Sciences, Centre for Ecological
Research Balaton Limnological Institute, Klebelsberg
Kuno u ´t 3, Tihany 8237, Hungary
e-mail: toth.viktor@okologia.mta.hu
123
Hydrobiologia
DOI 10.1007/s10750-016-2684-z