© Color. Technol., 121 (2005) 291 Web ref: 20050601
Coloration
Technology
Society of Dyers and Colourists
Dyeing behaviour of cotton fabric
bioscoured with pectate lyase and
polygalacturonase
M Calafell,
a,
* B Klug-Santner,
b
G Guebitz
b
and P Garriga
a
a
Chemical Engineering Department, UPC-ETSEIT, Colom 11, 08222 Terrassa, Spain
Email: margarita.calafell@upc.es
b
Institute of Environmental Biotechnology, Graz University of Technology, Austria
Received: 12 May 2005; Accepted: 16 August 2005
Bioscouring of cotton is normally carried out using pectinases having pectate lyase activity. The present
study has examined the influence of pectate lyase and hydrolase on the surface of cotton fibre. Dye
uptake by cotton scoured with polygalacturonase is found to be much lower than that scoured with
pectate lyase, Pectate lyase gave better dye exhaustion at 90 °C. The difference in dye exhaustion after
scouring with sodium hydroxide, polygalacturonase or pectate lyase, may be due to variation in the
amount of surfactant retained by the fibre, as this hinders dye uptake. This was particularly the case with
polygalacturonase. Reduced numbers of free carboxy groups on the fibre surface could indicate lower
pectin content. On the other hand it could, however, be due to the esterification of carboxy groups in the
pectin by hydroxy groups in the surfactant.
Introduction
Bioscouring is a relatively new technique for removing
pectin, proteins and waxes from raw cotton by means of
enzymes. This technique has advantages compared with
chemical scouring using sodium hydroxide, including
energy savings and reduced salt concentration in the
wastewater. In addition, less damage is caused to the fibre
and as a result less reduction in fibre strength.
The use of pectinases in bioscouring offers advantages
over other enzymes, including cellulases, lipases and
proteases [1,2]. Most current bioscouring processes for
cotton employ alkaline pectinases with pectate lyase
activity [3,4]. On the other hand, a number of micro-
organisms are found in nature which produce pectinases
with activity in acid media. Pectinases of this type, for
example polygalacturonase, exhibit hydrolase activity and
are used widely in the food processing industry [5].
The fact that hydrolases are not generally used in the
textile field has been due to lack of understanding of
the pectin cleavage on bioscouring. Pectin acts as a net
which traps molecules such as glycoproteins, waxes and
xyloglucans in a non-covalent manner within the cuticle of
the cotton fibre [6]. The structural complexity of cotton and
its degree of polymerisation provide a protective barrier to
the cotton fibre. It is worth describing briefly the nature of
the structure since this is important in bioscouring.
Pectins are a family of complex polysaccharides, usually
amorphous, with a degree of polymerisation around 200–400
units [7]. Three pectic polysaccharides, homogalacturonan,
rhamnogalacturonan-I and substituted galacturonans, have
been isolated from the primary cell wall and their structures
characterised [8]. The main chain of homogalacturonan
contains 1,4-linked α-D-galactosyluronic acid residues.
Substituents are located at the C-2 or C-3 position of the
main chain. Substituents may be non-sugar (acetyl) or sugar-
like (D-galactose, D-xylose, L-arabinose and L-rhamnose).
In cotton fibres, 60–75% of the chains of polygalacturonic
acid can be methylated. Rhamnogalacturonan-I contains a
backbone of repeating disaccharides [-4)-α-D-galacturonic
acid-(1,2)-α-L-rhamnose-(1-]. The galacturonic acid residues
in the backbone can be O-acetylated at C-2 and/or C-3. [9].
Rhamnogalacturonan-I is solubilised from cell walls by
polygalacturonases [10]. Substituted galacturonans are a
diverse group of polysaccharides containing a backbone of
linear 1,4-linked α-D-galacturopyranosyluronic acid residues
[11]. The polymer is highly methylated in the main chain
of polygalacturonic acid and has a greater number of side-
chains [12]. Due to the vulnerability of these polymers to
enzymes, pectic enzymes can cause plant tissue maceration,
cell lysis and modification of cell wall structure [13].
Pectic enzymes are produced in large quantities by many
plant-associated micro-organisms. Pectinolytic micro-
organisms produce a battery of pectic enzymes differing
in substrate preference, reaction mechanism and mode of
action. Pectate lyase, pectin lyase and exo-polygalacturonate
lyase cleave by β -elimination and generate products
having 4,5-unsaturated residues at the non-reducing end
(A in Scheme 1). Polygalacturonase and exo-poly- α- D-
galacturonosidase cleave by hydrolysis (B in Scheme 1).
Pectinesterase de-esterifies pectin to pectate and methanol.
Pectate lyase, pectin lyase, and polygalacturonase cleave
internal glycosidic bonds and generate a series of oligomeric
products. exo-Polygalacturonate lyase and exo-poly- α-D-
galacturonosidase attack chain termini, releasing only
digalacturonates, and reduce the viscosity of solutions
containing pectic polymers more slowly than the endo-
attacking enzymes. Pectin lyase attacks pectin (poly-
methoxygalacturonide); the other enzymes attack pectate
(polygalacturonate) [13].
In the present study, two different commercial pectinase