THE FUNCTION OF TRIUMPH CEREMONIES
IN THE BLACK SWAN
by
KEN KRAAIJEVELD
1)
and RAOUL A. MULDER
2)
(Department of Zoology, University of Melbourne, Vic 3010, Australia)
(Acc. 7-XI-2001)
Summary
The function of triumph ceremonies was investigated in the socially monogamous black
swan, Cygnus atratus . Paired black swans regularly performed triumph ceremonies, most
of which were initiated by the male. We found no difference in the frequency with which
ceremonies were performed by pairs prior to the breeding season compared to pairs with
dependent cygnets. Pairs with more cygnets tended to perform fewer triumph ceremonies
and the number of triumphs performed by a pair did not correlate with the survival of their
cygnets. Feeding experiments showed that the frequency of triumph ceremonies performed
by pairs increased when many swans were in close proximity. Pairs also performed triumph
ceremonies in response to the playback of a triumph ceremony by another pair, but not in
response to playback of an advertisement call by an unpaired black swan. We suggest that
our results are compatible with the pair formation and collaborative threat hypotheses, but
not with the bond-maintenancehypothesis.
Keywords : triumph ceremony, black swan, threat display, cooperation, pair-bond mainte-
nance.
1)
Corresponding author: e-mail address: ken_kraaijeveld@hotmail.com
2)
We thank the members of the Ballarat Field Naturalists, in particular John and Elaine
Gregurke, Carol Hall, Brian, Kevin and Joan Andrews for their help in eld. The Ballarat
City Council and the Dept. of Natural Resources and Environment permitted us to conduct
this study at Lake Wendouree. Femmie Kraaijeveld-Smitkindly read through an early version
of the manuscript. Jerry Hogan and an anonymous referee provided helpful comments. This
research was supported by grants from the Holsworth Wildlife Research Fund and the Stuart
Leslie Bird Research Fund.
© Koninklijke Brill NV, Leiden, 2002 Behaviour 139, 45-54