American Journal of Epidemiology
Copyright © 1995 by The Johns Hopkins University School of Hygiene and Public Health
All rights reserved
Vol. 142, No. 12
Printed in U.S.A.
Estimated Timing of Mother-to-Child Human Immunodeficiency Virus Type 1
(HIV-1) Transmission by Use of a Markov Model
C. Rouzioux,
1
D. Costagliola,
2
M. Burgard,
1
S. Blanche,
3
M. J. Mayaux,
4
C. Griscelli,
3
A.-J. Valleron,
2
and
the HIV Infection in Newborns French Collaborative Study Group
5
It has been shown that mother-to-child human immunodeficiency virus type 1 (HIV-1) transmission can
occur both during pregnancy and at delivery, but the respective frequencies in these periods are unknown.
Moreover, it is difficult to determine the timing of mother-to-child HIV-1 transmission by direct sampling. The
use of an elaborate statistical method is therefore necessary. The authors studied 495 consecutive infants born
between May 1988 and August 1991 who were included, at birth, in the French Prospective Study on Pediatric
HIV Infection. At least one blood sample was obtained from every infant during the first 14 days of life. All
samples obtained within 3 months of birth were tested by at least two of the following methods: viral culture,
polymerase chain reaction (PCR), and antigenemia, as well as by Western blot test. Data for the 95 infected
infants (those seropositive at 18 months and those who died of HIV disease before this age), and who were
exclusively bottle-fed, were analyzed in a Markov model to estimate the timing of viral transmission, the time
from birth to the emergence of detectable virus, and the time from birth to seroconversion. The model
indicated that one-third of the infants were infected in utero, less than 2 months before delivery (95th
percentile). In the remaining 65% of cases (95% confidence interval (Cl) 22-92), the date of infection was
estimated as the day of birth. The estimated median period between birth and the emergence of viral markers
was 10 days (95% Cl 6-14) and the 95th percentile was estimated at 56 days. These results support the view
that HIV infection can be diagnosed during the first 3 months of life. The authors conclude that mother-to-child
HIV-1 transmission appears to occur late in pregnancy or at delivery. Am J Epidemiol 1995;142:1330-7.
cohort studies; HIV-1; Markov models; mother-to-child transmission; statistics
The mother-to-child human immunodeficiency vi-
rus type 1 (HIV-1) transmission rate has been esti-
Received for publication June 6, 1994, and in final form March
30, 1995.
Abbreviations: AIDS, acquired immunodeficiency syndrome;
anti-HIV-1, antibody to human immunodeficiency virus type 1;
HIV-1, human immunodeficiency virus type 1; PBMC, peripheral
blood mononuclear cells; PCR, polymerase chain reaction.
1
Laboratoire de Virologie, Hopital Necker-Enfants Malades, 149
rue de Sevres, 75015 Paris, France. (Correspondence to Prof. C.
Rouzioux at this address.)
2
B
3
E, INSERM U 263 et SC4, Universite Pierre et Marie Curie,
Faculte de Medecine Saint-Antoine, Paris, France.
3
Unite d'lmmunologie-Hematologie, Hopital Necker-Enfants
Malades, Paris, France.
4
INSERM U 292, Hopital du Kremlin-Bicetre, Le Kremlin-Bicetre,
France.
5
The following persons are participants in the HIV Infection in
Newborns French Collaborative Study: M. C. Allemon, D.
Armengaud, J. M. Babinet, P. Balde, F. Ben Fadel, R. Bensadoun,
D. Berterotiere, M. Boddaert, Y. Bompard, C. Botto, A. M. Brunner,
J. A. Cacault, C. Carlus-Moncomble, N. Ciraru-Vigneron, C. Cohen,
A. M. Colin-Gorki, C. Courpotin, C. Crumiere, M. C. Dallot, M.
Dandine, A. De Crepy, M. Debons, M. Debre, M. F. Denavit, A.
Devidas, M. Dumontel, G. Firtion, C. Floch, C. Francoual, J. Furioli,
A. Gantzer, F. Granier, F. Guillot, B. Heller, C. Huraux-Rendu, P.
Labrune, E. Lachassine, A. Lacroix, M. Lanza, H. Lajalle, B. Le
Lorier, A. Leblanc, F. Lebrun, C. Lejeune, J. Mamou, A. May, F.
Mazi, A. Meyer, G. Mouchnino, P. Narcy, G. Noseda, C. Pascal, B.
mated at between 10 and 39 percent in epidemiologic
studies (1). The timing of transmission is unknown,
yet it is crucial to know for effective prevention.
Lentiviruses, including HIV-1, replicate and can be
detected in the blood throughout the course of the
infection; HIV-1 is continuously present in the form of
complete infectious virions in the plasma and as pro-
viruses integrated in host lymphocyte DNA. This
means that the fetus of an infected woman is, in
theory, continuously exposed to the virus. Early trans-
mission has been suggested by reports of HIV-1 DNA
detection in aborted fetuses and fetal tissues taken
during the first 6 months of pregnancy (2, 3), while
other studies have pointed to later transmission in
utero or during delivery (4, 5). HIV-1 is detectable at
birth in only about 40 percent of infected infants (6,7),
suggesting that infection occurs close to or during
delivery in the remaining cases. Firstborn twins are
more frequently infected than their siblings, which
Pautard, M. Robin, M. Ronzier, J. L. Ropert, M. C. Rousset, A.
Saillant, G. Scart, H. Seaume, D. Seguy, P. Talon, M. Tardieu, J.
Terris, L. Valdes, F. Veber, J. Vedrenne, M. Vial, A. Vinas, N. Vincent,
and P. Wipff.
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