Long-term potentiation saturation in chronic cerebral
hypoperfusion
Lali H.S. Sekhon 1 MB BS PhD, lan Spence 2 BSc PhD, Michael K. Morgan I MD FRACS,
Neville C. Weber 3 MSc PhD
1Department of Surgery DO6, The Universityof Sydney, Sydney,NSW 2006, Australia
2Department of Pharmacology DO5, The Universityof Sydney,Sydney,NSW 2006, Australia
3School of Mathematics and Statistics FO7, The Universityof Sydney, Sydney,NSW 2006, Australia
Chronic reductions in cerebral blood flow (CBF) of between 25 and 50%, in the absence of cerebral infarction, lead
to impairments in hippocampal in vitro long-term potentiation (LTP). This study set out to explore some of the
properties of this impairment of LTP. LTP is an electrophysiological property known to occur in the hippocampus, a
region known to be exquisitely sensitive to hypoxic or ischemic insults. Thus, assessing LTP is a novel way of assessing
the effects of subtle ischemic insults. Five male Sprague-Dawley rats had arteriovenous fistulae created surgically in
the neck to induce a state of chronic cerebral hypoperfnsion (CCH) with the features described above. Five rats were
used as age-matched controls. Twenty-six weeks after fistula formation, the animals were prepared for in vitro
hippocampal recording in a submerged tissue bath. Extracellular field potentials were recorded at the Schaffer
collateral-CA1 region, with a stimulus intensity that achieved a population spike amplitude of 1 mV. After tetanic
stimulation, the frequency and magnitude of LTP was compared between control and fistula animals. All animals in
both these groups demonstrated LTP in contradistinction to our previous work where LTP was impaired in fistula
animals when a higher intensity of stimulation was used. This indicates that the structures that are associated with the
initiation and maintenance of LTP (most probably the ischemia-sensitive CA1 pyramidal cells) are saturated as the
stimulus intensity is increased. Thus, at this lower intensity of stimulation LTP is preserved in the fistula animals, but
found to be impaired as the stimulus intensity is increased. Consequently, this study provides further information on
this newly identified subtype of chronic cerebral ischemia which, in time, after further studies in humans, may help
to redefine therapeutic indicators for the management of cerebral arteriovenous malformation and severe
cerebrovascular disease.
Journal of Clinical Neuroscience 1998, 5 (3) : 323-328 © Harcourt Brace & Co. Ltd 1998
Keywords" arteriovenous malformation, brain slices, electrophysiology, hippocampus, ischemia, long-term potentiation
Introduction
Chronic reductions in cerebral blood flow (CBF) can
occur in many clinical situations, best typified by the
reduction in CBF that occurs as a result of venous hyper-
tension and cerebrovascular 'steal '1 in brain parenchyma
adjacent to cerebral arteriovenous malformations (AVMs).
Numerous clinical reports have described patients who
have had neurological dysfunction thought attributable to
the shunting of blood from adjacent neuronal structures
with cerebral AVMs, in the absence of cerebral infarc-
tion3 -7 Brown et al a noted that 9 of 135 patients in their
study with AVMs had a functional decline over the 4 years
of follow-up, translating to a functional decline in 1.6% of
patients with AVMs per year. This decline was not
attributed to hemorrhage and presumably would, for the
most part, be attributable to the effects of the preferential
diversion of blood flow through the low resistance vessels
of an AVM, creating chronic cerebral hypoperfusion
(CCH) in brain regions adjacent to the AVM. CCH can
also occur in severe bilateral carotid artery disease, and
may be responsible for subtle personality and intellect
degradations that occur in these patients without evidence
of infarction on computed tomography (CT) scanning.
Up until the past few years it was thought that the
thresholds for neuronal dysfunction described for acute
ischemic insults were applicable when hypoperfusion was
sustained for a prolonged duration? -n More recently,
however, we have shown that there is a new subtype of
chronic cerebral ischemia which does not obey the
classical thresholds for acute ischemic insults. Chronic
reductions in CBF of between 25 and 50%, in the absence
of cerebral infarction impair neuronal function, 12 and
alter structure 13a4 with these changes present after
approximately 6 months of constant hypoperfusion. The
changes are not present after only 10 weeks of this degree
of hypoperfusion] 5 which implies that whatever mecha-
nisms are responsible for these changes take time to
manifest, with a maturation process initiated prior to the
induced changes becoming overt.
J. Clin. Neuroscience Volume 5 Number 3 July 1998 323