Detection of Amplifiable mRNA Extracellular to
Insulin-Producing Cells: Potential for Predicting
Beta Cell Mass and Function
Sweta Rani, Martin Clynes, and Lorraine O’Driscoll
*
Background: Detecting extracellular nucleic acids in the
serum/plasma of cancer patients may help in cancer diag-
nosis. We investigated whether extracellular mRNAs are
reproducibly detectable in conditioned medium (CM)
from insulin-producing cell cultures and if their presence
and amounts are indicative of cell number and/or function.
Methods: We isolated mRNA from medium condi-
tioned by the culture of several insulin-producing cell
types: MIN6(L) (glucose-responsive), MIN6(H) (glucose-
nonresponsive), and MIN6 B1 murine beta cells and
monkey kidney fibroblast cells engineered to produce
human preproinsulin (PPI) (Vero-PPI). We used reverse
transcription–PCR analyses to evaluate the occurrence
of several mRNAs and investigated whether the pres-
ence and amounts of the various extracellular mRNAs
are associated with cell mass and/or function.
Results: Reproducible amplification of mRNAs en-
coded by Pdx1, Npy, Egr1, Pld1, Chgb, Ins1, Ins2, and
Actb from MIN6(L), MIN6(H), and MIN6 B1 cells and
their CM suggests that beta cells transcribe and release
these mRNAs into their culture environment. Similarly,
PPI mRNA was detected in samples of Vero-PPI cells
and CM. The amounts of some mRNAs reflected the
numbers and functional status (i.e., glucose responsive-
ness vs nonresponsiveness) of the cells conditioning the
medium. Although Pax4 mRNA was detected in the
MIN6 B1 cell line, the fact that this transcript was not
amplifiable from the corresponding CM suggested that
mRNA release was selective.
Conclusion: mRNAs may be secreted from insulin-
producing cells, are reproducibly detected in the extra-
cellular environment, and may have potential as extra-
cellular biomarkers for assessing beta cell mass and
function.
© 2007 American Association for Clinical Chemistry
Diabetes currently affects 246 million people worldwide
and will affect 380 million by 2025 (1). Type 1 diabetes is
characterized by autoimmune destruction of the insulin-
producing beta cells in the islet of Langerhans of the
pancreas, whereas type 2 diabetes is associated with
reduced beta cell mass and function, as well as with
insulin resistance. No panel of circulating biomarkers of
beta cell mass and/or function has been reported. Because
type 1 diabetes is usually diagnosed after a loss of
70%– 80% of the insulin-secreting beta cells in the pan-
creas (2), detecting an association of gene expression in
the circulating serum with beta cell loss and/or function
may aid in early disease detection.
Nucleic acids were first detected in plasma almost 60
years ago (3), and with the recent advances in sensitive
and specific analytical techniques, the detection of nucleic
acids now has potential as a tool for early diagnosis of
disease. Extracellular RNA has been detected in the
plasma and serum of patients with various forms of
cancer (4–6), DNA and mRNA of fetal origin have been
discovered in the plasma of pregnant women (7–9 ), and
significantly increased concentrations of rhodopsin
mRNA have been found in the peripheral blood of
individuals with diabetes (10 ). Although RNA biomark-
ers indicative of pancreas beta cell mass and function may
circulate in diabetic patients, no studies have yet investi-
gated this possibility. We previously reported methods
for detecting mRNAs extracellular to cancer cells (11 ). In
an extension of this approach, we investigated whether
extracellular mRNAs are reproducibly detectable in beta
National Institute for Cellular Biotechnology, Dublin City University,
Dublin 9, Ireland.
* Address correspondence to this author at: National Institute for Cellular
Biotechnology, Dublin City University, Dublin 9, Ireland. Fax 353-1-7005484;
e-mail Lorraine.ODriscoll@dcu.ie.
Received February 21, 2007; accepted August 7, 2007.
Previously published online at DOI: 10.1373/clinchem.2007.087973
Clinical Chemistry 53:11
1936 –1944 (2007)
Endocrinology and
Metabolism
1936