446 Accepted by H. Chiba: 16 Jul. 2015; published: 11 Aug. 2015 ZOOTAXA ISSN 1175-5326 (print edition) ISSN 1175-5334 (online edition) Copyright © 2015 Magnolia Press Zootaxa 3999 (3): 446–450 www.mapress.com/zootaxa/ Correspondence http://dx.doi.org/10.11646/zootaxa.3999.3.10 http://zoobank.org/urn:lsid:zoobank.org:pub:EF30014F-DCF5-4457-96AA-A02CE48B4748 Tosta Evans, 1953 (Lepidoptera: Hesperiidae): a new species from northeast Brazil, first description of the female of Tosta tosta Evans, 1953, and placement of the genus within Achlyodini RICARDO RUSSO SIEWERT 1 , OLAF HERMANN HENDRIK MIELKE 2 & MIRNA MARTINS CASAGRANDE 3 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, P.O. Box 19.020, ZIP Code 81.531-980, Curitiba, Paraná, Brazil. E-mail: 1 ricardo.siewert@gmail.com; 2 omhesp@ufpr.br; 3 mibras@ufpr.br Abstract. A new species of Tosta Evans, 1953 from northeast Brazil is described: T. pseudospeculum Siewert, Mielke & Casagrande, sp. nov. Additionally, the female of Tosta tosta Evans, 1953, the type species of the genus, is described for the first time. Illustrations of adults and male and female genitalia for all studied species are provided, as well as a discussion of the placement of the genus within the tribe Achlyodini. Keywords: Achlyodini, butterfly, Erynnini, skippers, taxonomy Resumo. Uma nova espécie de Tosta Evans, 1953 do nordeste do Brasil é descrita: T. pseudospeculum Siewert, Mielke & Casagrande, sp. nov. Adicionalmente, a fêmea de Tosta tosta Evans, 1953, a espécie tipo do gênero, é descrita pela primeira vez. Ilustrações dos adultos e das genitálias masculinas e femininas das espécies estudadas são fornecidas, bem como uma discussão sobre a transferência de Tosta para a tribo Achlyodini. Palavras chave: Achlyodini, borboleta, Erynnini, hesperídeos, taxonomia Introduction Tosta Evans, 1953 (Hesperiidae: Pyrginae) is a Neotropical genus of skippers currently including eight species (Mielke 2004, 2005, Steinhauser 2007). Evans (1953) described the genus to incorporate five species allied to Anastrus Hübner, [1824] and Achlyodes Hübner, [1819], which are now located in Erynnini and Achlyodini, respectively. In the phylogenetic study proposed by Warren et al. (2008, 2009), Tosta was assigned to Erynnini. However, Tosta seems to be polyphyletic, and some species that are currently included in the genus probably belong to another genus or even another tribe (Warren et al. 2009). Other aspect that difficults the study of Tosta is the supposed scarcity of the specimens in collections, and until now, all females remains unknown. This paper aims to describe a new species of Tosta from northeast Brazil. Additionally, the female of Tosta tosta Evans, 1953 (type species of the genus) is described for the first time. Detailed illustrations of the adults, male and female genitalia, diagnosis, distributional information and taxonomic comments are provided for all studied species. Material and methods The specimens studied were dissected using standard techniques for Lepidoptera. Dissected specimens are marked with an asterisk after the voucher number in the examined material section. All specimens are deposited at Coleção Entomológica Padre Jesus Santiago Moure, Universidade Federal do Paraná, Curitiba, Paraná, Brazil (DZUP). Results Tosta pseudospeculum Siewert, Mielke & Casagrande sp. nov. (Figs 1–4, 9–13, 19–20)