Citation: Ponizovsky AM, Barshtein G, Nechamkin Y, Lecht S, Bergelson LD. Differences in the Lipid Domain Organization of Erythrocyte Membranes in
Patients with Schizophrenia. J Neurol Psychol. 2015;3(1): 6.
J Neurol Psychol
January 2015 Vol.:3, Issue:1
© All rights are reserved by Ponizovsky et al.
Differences in the Lipid Domain
Organization of Erythrocyte
Membranes in Patients with
Schizophrenia
Ke ywords: Erythro c yte me mb ra ne ; Lip id d o ma ins; Fluo re sc e nc e
me a sure me nts; Sc hizo p hre nia ; Symp to ma to lo g y
Abstra c t
G ra d ua lly a c c umula ting e vid e nc e ind ic a te s tha t sc hizo p hre nia
ma y b e a c c o mp a nie d b y a lte ra tio ns o f me mb ra ne p ho sp ho lip id s,
ho we ve r kno wle d g e a b o ut ho w it a ffe c ts lip id d o ma in o rg a niza tio n
of membrane is lacking. We compared the membrane lipid domain
o rg a niza tio n o f re d b lo o d c e ll (RBC ) me mb ra ne s fro m two g ro up s o f
p a tie nts with sc hizo p hre nia se le c te d b y the ir hig h p o sitive synd ro me
(PS) o r ne g a tive synd ro me (NS) sc o re s o n the Po sitive a nd Ne g a tive
Synd ro me Sc a le , a nd c o ntro l sub je c ts witho ut kno wn p syc hia tric histo ry.
Diffe re nc e s in the surfa c e d istrib utio n o f the me mb ra ne p ho sp ho lip id s
were elucidated by: 1) registering the luorescence excitation spectra
o f the p o la rity-se nsitive lip id p ro b e La urd a n inc o rp o ra te d into the RBC
me mb ra ne , 2) me a suring the re so na nc e e ne rg y tra nsfe r (RET) fro m
tryptophan to luorescent analogs of the main RBC phospholipids
p ho sp ha tid ylc ho line a nd sp hing o mye lin, a nd 3) d e te rmining the
c ha ng e s in RET fro m tryp to p ha n to the sp hing o mye lin p ro b e ind uc e d
b y p ro sta g la nd in E1. The d a ta o b ta ine d with RBC s fro m e ithe r no rma l
subjects or PS patients were largely similar, but differed signiicantly
fro m tho se o b ta ine d with RBC s o f p a tie nts with NS sc hizo p hre nia . The
re sults sup p o rt va lid ity o f the c linic a l d istinc tio n o f sc hizo p hre nia into
p o sitive a nd ne g a tive sub typ e s.
Introduction
Most approaches to subtyping schizophrenia have made use of
the concept of positive and negative syndromes (PS and NS). PS are
characterized by hallucinations, delusions and thought disorders
whereas apathy, emotional blunting and psychomotor retardation
are regarded as NS [1]. Although the positive-negative dichotomy is
believed by some authors to be a simpliication [2-4], it has important
clinical and prognostic signiicance. A number of recent investigations
have revealed that the positive and negative syndromes are associated
with diferent biochemical changes. Particularly, previous studies
have detected certain speciic lipid aberrations in red blood cells
(RBC) from psychiatric patients with predominantly negative
symptoms [5-8]. Recently, we reported on noticeable increases in
both the sphingomyelin level and the aggregability of RBCs from
patients with schizophrenia with mainly NS in comparison to
those from patients with schizophrenia with mainly PS and normal
controls [9,10]. Previously, RBCs from patients with schizophrenia
were found to be enriched also in cholesterol and glycolipids [11-
13] and their membranes were reported to be less “luid” than those
of normal erythrocytes [11,13]. However the concept of membrane
luidity depends on the assumption that the hydrophobic region
of the membrane is structurally and dynamically homogeneous,
whereas in eukaryotic cells including human and rabbit erythrocytes
[14-18], the surface lipids appear to distribute unevenly between
Alexander M. Ponizovsky
1
*, Gregory Barshtein
2
,
Yakov Nechamkin
3
, Shimon Lecht
4
and Lev D.
Bergelson
4
1
Research Unit, Mental Health Services, Ministry of Health,
Jerusalem, Israel
2
Department of Biochemistry, Hadassah Medical School, The
Hebrew University of Jerusalem, Israel
3
Sha’ar Menashe Mental Health Center, Hadera, Israel
4
Biological Membrane Group, School of Pharmacy, the Hebrew
University of Jerusalem, Israel
*Address for Correspondence
Alexander M. Ponizovsky, MD, PhD, Research Unit, Mental Health
Services, Ministry of Health, 39 Yirmiyahu St., PO Box 1176, Jerusalem
9446724, Israel, E-mail: alexpon8@gmail.com
Submission: 24 November 2014
Accepted: 22 December 2014
Published: 29 December 2014
Research Article Open Access
Journal of
Neurology and
Psychology
Avens Publishing Group
Invi ting Innovations
lateral micro-domains. Since the activity of membrane receptors and
membrane bound enzymes depends on the physical state of their
nearest environment, local properties pertaining to lipid domains in
the immediate neighborhood of membrane proteins may be more
relevant than bulk lipid properties. Nevertheless the connection
between lipid domain structure and pathophysiological events is a
new, still poorly explored ield of clinical biochemistry. In recent years
it has been found that in many cells sphingolipids and cholesterol
cluster together forming movable domains or rats, which are believed
to be involved in signal transduction and cell-cell interactions [19].
In view of the unusually high sphingomyelin (SM) levels in RBC
from patients with NS schizophrenia [9] we therefore thought it
of interest to obtain more information about the lipid distribution
in the RBC membrane, particularly, the lateral segregation of its
two main phospholipid species, SM and phosphatidylcholine (PC)
in membranes of RBCs isolated from the blood of patients with
schizophrenia with deined clinical symptom scale scores. In this
study we attempted to detect diferences in the lipid domain structure
of erythrocytes from normal subjects and subgroups of patients with
patients with PS or NS schizophreniaby studying excitation spectra
of the polarity-sensitive luorescent membrane probe Laurdan, as
well as by measuring the luorescence resonance energy transfer
(RET) from protein tryptophans to luorescent analogs of SM and PC
incorporated into the erythrocyte membrane.
Some earlier studies of polyunsaturated fatty acids, prostaglandin
synthesis and prostaglandin receptor activity suggested a role for
prostaglandins in the pathophysiology of schizophrenia (reviewed in
[20]). Because red blood cells are known to be extremely sensitive
to prostaglandin E
1
(PGE
1
) [15,21-26] (see also [21,22] for reviews
of the older literature), we here use the multilipid probe approach
also to detect possible diferences between the response to PGE
1
of the
phospholipid domain organization in the membranes of normal red
cells and those isolated from the blood of patients with predominantly
NS schizophrenia.