Research Article
Floristic Composition, Structure, and Species Associations of
Dry Miombo Woodland in Tanzania
Ezekiel Edward Mwakalukwa,
1,2
Henrik Meilby,
1
and Thorsten Treue
1
1
Department of Food and Resource Economics, Faculty of Science, University of Copenhagen, Rolighedsvej 23,
1958 Frederiksberg C, Denmark
2
Department of Forest Biology, Faculty of Forestry and Nature Conservation, Sokoine University of Agriculture,
P.O. Box 3010, Chuo Kikuu, Morogoro, Tanzania
Correspondence should be addressed to Ezekiel Edward Mwakalukwa; ezedwa@yahoo.com
Received 20 December 2013; Accepted 4 February 2014; Published 8 May 2014
Academic Editors: M. Drielsma, H. Ford, M. Tigabu, and A. Vi˜ na
Copyright © 2014 Ezekiel Edward Mwakalukwa et al. his is an open access article distributed under the Creative Commons
Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is
properly cited.
For the majority of forest reserves in Tanzania, biodiversity is poorly documented. his study was conducted to assess species
richness (woody species), diversity, and forest structure and to examine relationships between species occurrence and topographic
and edaphic factors in the Gangalamtumba Village Land Forest Reserve, a dry Miombo woodland area in Tanzania. A total of 35
nested circular plots with radii of 5, 15, and 20m were used to collect data on woody species and soil samples across the 6,065 ha
community-managed forest reserve. Stumps were measured 20cm above ground. A total of 88 species belonging to 29 families
were identiied. Generally forest structure parameters and diversity indices indicated the forest to be in a good condition and have
high species richness and diversity. Vegetation analysis revealed four communities of which two were dominated by the family
Caesalpiniaceae, indicating large variation of site conditions and possible disturbances in the study area. he high level of diversity
of woody species and the high basal area and volume indicate that the forest is in good condition, but the efect of anthropogenic
activities is evident and stresses the need for proper management to maintain or enhance the present species diversity.
1. Introduction
Miombo woodland is the most widespread and dominant dry
forest formation in Eastern, Central, and Southern Africa. It
is characterized by an abundance of tree species in the legume
subfamily Caesalpinoideae, including the three dominant
genera of Brachystegia, Julbernardia, and Isoberlinia [1, 2].
Covering an area of about 3.6 million km
2
, miombo wood-
land supports the livelihoods of more than 100 million rural
and urban dwellers by providing a wide range of products
such as irewood, charcoal, timber, and forage and services
such as soil conservation and water catchment [3–5]. How-
ever, due to the rapid population growth and the high level of
poverty across the Miombo region, the human pressure on its
woodlands has steadily increased over the last decades, lead-
ing to increasing deforestation and forest degradation [6–8].
he efects of increasing rates of deforestation and forest
degradation on biodiversity in developing countries have
been thoroughly studied [9–12]. Habitat loss due to deforesta-
tion reduces not only the number of species in the ecosystem
but also the number and extent of places where species
coexist. Activities such as charcoal production, irewood col-
lection for subsistence use and for tobacco curing, conversion
of woodlands to farmland, and seasonal forest ires are among
the major drivers of deforestation and forest degradation in
the Miombo region [13–17]. It is estimated that 1.4 million
ha of woodlands is lost annually in the countries where
Miombo woodlands dominate, leading to a loss of carbon
stocks, biodiversity, and, through soil degradation, loss of
plant nutrients [4, 5]. Syampungani et al. [5, p. 151] stated that
“loss of biodiversity and extinction of most of the woodland
resources are imminent if the current intensive exploitation of
Miombo resources continues unchecked.” More speciically,
FAO (2000a, cited by Syampungani et al. [5]) reported that
191 tree species in the Miombo ecoregion are endangered due
Hindawi Publishing Corporation
ISRN Biodiversity
Volume 2014, Article ID 153278, 15 pages
http://dx.doi.org/10.1155/2014/153278