Journal of Paleolimnology 14: 49--67. 1995. 49
© 1995 Kluwer Academic Publisher.f. Printed in Belgium.
An expanded weighted-averaging model for inferring past total phosphorus
concentrations from diatom assemblages in eutrophic British Columbia
(Canada) lakes
Euan D. Reavie', Roland I. Hall',2 & John P. Smol'
1 Paleoecological Environmental Assessment & Research lLJb (PEARL), Department of Biology, Queen j.
University, Kingston, Ontario, Canada, K7L 3N6
2present address: Ontario Ministry of Environment & Energy, Science & Technology Branch, Aquatic Science
Section, Bellwood Acres Rd., P. 0. Box 39, Dorset, Ontario, Canada, POA J EO
Received 12 August 1994; accepted 19 December 1994
Key words: diatoms, eutrophication, lake management. paleolimnology, British Co lumbia, lakes. phosphorus,
training sets
Abstract
Eighteen lakes were added to a published training set of 46 British Columbia (BC) lakes in order to expand the
original range of total phosphorus (TP) concentrations. Canonical corresponden ce analysis (CCA) was used to
analyze the relationship between diatom assemblages and environmental variables. Specific conductivity and {TP]
each exp lained significant (PSO.05) directions of variance in the distribution of the diatoms. The relationship
between diatom assemblages and [TP] was sufficiently strong to warrant the development of a
(WA) regression and calibration model that can be used to inf er past trophic status from fossil diatom assemblages.
The relationship between observed and inferred [TP] was not improved by the addition of more eutrophic lakes,
however the [TP] range and the number of taxa used in the transfer function are now superior to the original model.
Diatom species assemblages changed very little in lakes with TP concentrations greater than 85 j.Lg 1-1, so we
document the development of a model co ntaining lakes with p,g 1- 1. The updated model uses 59 training
lakes and covers a range of species optima from 6 to 41.9 J.Lg 1-1 TP, and a total of 150 diatom taxa.
The updated inference model provided a more realistic reconstruction of the anthropogenic history of a highly
eutrophic BC lake. The model can now be used to infer past nutrient conditions in other Be lakes in order to assess
changes in trophic status.
Introduction
Water quality has declined in many western Canadi-
an lakes as a result of anthropogenic nutrient loading.
These lakes have often been subject to perturbations
since European settlement began in the 1800s, and
in some areas, Indian settlements may have affected
water quality well before this time (Woolliams, 1979).
However, water quality information measured before
1950 is scarce or lacking for most systems, and realistic
estimates of previous ecological conditions are often
only available in the form of proxy data from the sed-
iment record. Paleolimnological approaches can
vide information on the causes, timing and extent of
ecological disturbances. Such data are often necessary
for proper management decisions (Smol, 1992). More-
over, with knowledge of the pre-disturbance state,
istic restoration goals can be estab li shed.
The siliceous remains (valves) of diatoms have
been widely used as biomarkers to elucidate environ-
mental conditions in lakes (e.g. Dixit et aI., 1992).
Recently, ecologicall y relevant statistical methods
have been developed for inferring environmental
ditions from diatom assemblages. These methods are
based on multivariate ordination and weighted
aging rNA) regression (that effectively estimates the
optima and tolerances of several species) and calibra-
tion (Birks et at., 1990a). Ecological parameters of the