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0090-2977/14/4604-0376 © 2014 Springer Science+Business Media New York
Neurophysiology, Vol. 46, No. 4, August, 2014
Do Results of the Open Field Test Depend on the Arena
Shape?
S. V. Grabovskaya
1
and Yu. T. Salyha
1
Received May 05, 2014
The open field test (OFT) is one of the main methods used for the estimation of behavioral indices and
emotionality levels in animals. This test is widely used in toxicological and pharmacological studies.
The question of whether the arena shape influences the results of testing remained open. This is why
we compared the results of the mentioned test using one and the same species of laboratory animals
(rats), identical conditions of testing, and arenas of nearly the same area but of a round or square shape.
Significant differences were found in only two examined parameters, frequency of realization of the
burrow reflex and intensity of “internal” horizontal motor activity; other indices showed nearly no
considerable differences. Thus, it can be concluded that the results of behavioral studies carried out
under the above two conditions are rather similar to each other in their adequacy.
Keywords: open field test, behavioral indices, arena shape, rats.
Institute of Biology of Animals of the National Academy of Sciences of
Ukraine, L’viv, Ukraine
Correspondence should be addressed to Yu. T. Salyha
(yursalyha@yahoo.com)
INTRODUCTION
The open field test (OFT) is one of the most
popular and extensively known methods in the
investigation of animal behavior. It was introduced
into the scientific practice in 1932 by Hall [1].
This test is based on the natural drive of animals,
rodents in particular, to examine a novel territory;
the respective behavioral phenomena are dissimilar
in animals with different levels of emotionality [2].
Most frequently, the OFT is used in investigations
of behavioral phenomena in mice [3, 4] or rats; there
are also studies carried out on gerbils [5], vole mice
[6], hamsters [7], dogs, primates, fish [8], and even
humans (10-month-old or younger children) [9].
In the course of testing, the animals are placed
in a square or round arena open on top and limited
by walls from the sides. Rectangular arenas
with different ratios of the side lengths are used
considerably more rarely. Materials for arenas used
in different studies vary (plastics, wood, metal,
rubber, or even glass); colors of the walls also can
be different (from white to black) [9]. The floor
of the arena is divided into squares (in the case of
square or rectangular arenas) or into segments (in
round arenas). As a rule, the internal space of the
arena is empty; some authors, however, place in this
space some additional objects (e.g., mirrors, pieces
of food, flashing sources of light, other animals in
cages, etc.). The testing period for one animal is
usually 3 to 5 min long; this period, however, can
be prolonged to one hour [10] or even longer.
Observation of the animal’s behavior is carried
out; indices of the latter are recorded immediately
in the course of the testing period, or behavior
is videorecorded for off-line analysis. The main
examined parameters are the following: (i) horizontal
motor activity (number of the squares or segments
crossed by the animal within the observation period),
this index most closely resembles the intensity of
locomotion; (ii) vertical motor activity (number of
the vertical risings on the hindlimbs); (iii) number
of defecations; (iv) number of urinations, and (v)
number of grooming episodes. In addition, hori-
zontal motor activity is frequently divided into
external (crossing the squares or segments near
the wall of the arena) and internal (movement of
the animal toward the central part of the arena and
crossing the central squares of segments). Grooming
can be classified as short-lasting (two of three
movements of the forelimbs near the head) or long-
lasting (more movements combined with cleaning
of other parts of the body). The so-called burrow
reflex is interpreted as one of the indices of research
activity; this is sniffing of burrow-like openings in
the floor and attempts to enter them in the case of
sufficient dimensions. Some authors divide vertical
motor activity into a free one (risings on the hindlimbs
with no support) and that manifested near the wall
(with a support) [1]. There are studies where some